Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Wayne R. Ferren, Jr.

Annual, perennial herb, shrub, glabrous or hairy
Stem prostrate to erect
Leaves simple, generally basal and cauline, alternate, opposite, or whorled, rarely fleshy; stipules 0, conspicuous, or small and deciduous
Inflorescence: cyme, cluster, or flower solitary, axillary
Flower generally bisexual, small, radial; calyx persistent, sepals 4–5, generally free; corolla 0 or small; stamens 5–10, sometimes petal-like, attached to hypanthium, filaments free or fused at base; nectary a ring; ovary superior, chambers 1–10, placentas generally axile, styles 1 or 3–5, generally free
Fruit: generally capsule, generally loculicidal
Seeds 1 or more per chamber, sometimes with arils
Genera in family: 14 genera, 95 species: generally tropical, subtropical, especially Africa.


Annual, generally hairy
Stem prostrate to ascending, branched from base
Leaves alternate or appearing whorled, entire or toothed, petioled, unequal; stipules 0
Inflorescence: cluster head-like, flowers generally 5–10; pedicel short
Flower bell-shaped; sepals 5, margins scarious; petals 0; stamens 3–20, free or fused in groups, outer sterile, filaments short, slender; ovary chambers 3–5, style short, stigmas 3–5
Fruit ovoid
Seeds many, minute, smooth or tubercled; aril coiled around seed
Species in genus: 12 species: tropical, subtropical
Etymology: (Greek: sweet juice)


G. lotoides L.

Plant hairy; hairs generally forked or stellate
Stem 1–3 dm
Leaves entire; petiole ± = blade; blade 0.5–3 cm, obovate to round, ± gray-green, base tapered, tip generally abruptly acute, veins generally prominent
Inflorescence: flower ± sessile
Flower: sepals 4–7 mm, oblong or lanceolate, acute, keeled, tomentose
Fruit 4 mm
Seed 0.6 mm, tubercled, blackish brown
Ecology: Uncommon. Moist or seasonally dry margins of wetlands
Elevation: < 1000 m.
Bioregional distribution: North Coast Ranges, Great Central Valley, San Francisco Bay Area, Outer South Coast Ranges, South Coast, Western Transverse Ranges, San Gabriel Mountains, Peninsular Ranges
Distribution outside California: to Alaska, se US, etc.; native to Europe
G. radiatus (Ruiz & Pav.) Rohrb., native to tropical Am., is reported from ScV, PR: leaf green, sparsely hairy, seed 0.4 mm, generally shiny red-brown.

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