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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Dieter H. Wilken, except as specifed

Annual, perennial herb, shrub, glabrous to hairy, generally aromatic
Stems generally erect, generally 4-angled
Leaves generally simple to deeply lobed, opposite, generally gland-dotted
Inflorescence: cyme, generally clustered around stem, head-like, separated by evident internodes (terminal in Monardella ) or collectively crowded, spike-like to panicle-like (sometimes raceme or flowers 2–12); subtended by leaves or bracts; flowers sessile or pedicelled
Flower generally bisexual; calyx generally 5-lobed, radial to bilateral; corolla generally bilateral, 1–2-lipped, upper lip entire or 2-lobed, ± flat to hood-like, sometimes 0, lower lip generally 3-lobed; stamens generally 4, generally exserted, paired, pairs unequal, sometimes 2, staminodes 2 or 0; ovary superior, generally 4-lobed to base, chambers 2, ovules 2 per chamber, style 1, arising from center at junction of lobes, stigmas generally 2
Fruit: nutlets 4, generally ovoid to oblong, smooth
Genera in family: ± 200 genera, 5500 species: worldwide. Many cultivated for herbs, oils (Lavandula , lavender; Mentha , mint; Ocimum , basil; Rosmarinus , rosemary; Thymus , thyme), some cultivated as ornamental (in CA Cedronella , Leonotis , Phlomis )
Reference: [Cantino & Sanders 1986 Syst Bot 11:163–185]



Perennial from rhizomes, glabrous or hairy
Stems erect, branched or not
Leaves short-petioled to sessile, generally ovate to lanceolate; margin toothed to deeply lobed or cut below middle
Inflorescences axillary, each head-like, subtended by leaves
Flower: calyx generally 5-lobed, lobes ± equal, obtuse to short-awned; corolla slightly bilateral, not 2-lipped, generally 4-lobed, lobes ± unequal, odd lobe notched or entire; stamens 2, exserted, staminodes 2, minute, club-shaped; style exserted
Fruit: nutlets ± compressed, edge corky-thickened, truncate or rounded
Species in genus: 14 species: temp North America, Eurasia, 1 species in Australia
Etymology: (Greek: wolf foot, from French common name)
Reference: [Henderson 1962 Amer Midl Naturalist 68:95–135]


L. americanus W.C. Barton

Rhizomes ± slender, not thickened at tip
Stems erect, 2–8 dm, generally glabrous; nodes short-hairy
Leaf 2–8(10) cm, short-petioled, oblong to lanceolate, lobed to cut especially in lower half, glabrous to puberulent on veins
Flower: calyx lobes awl-like, short-awned; corolla 2–3 mm, ± = calyx, white
Fruit: nutlet 1–1.5 mm, top rounded, smooth
Chromosomes: 2n=22
Ecology: Moist areas, marshes, streambanks
Elevation: < 1000 m.
Bioregional distribution: California Floristic Province, East of Sierra Nevada
Distribution outside California: to British Columbia, e N.America
Flowering time: Aug–Sep

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