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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

KRAMERIACEAE

RHATANY FAMILY

Beryl B. Simpson

Perennial, shrub, green root-parasite
Stem prostrate to erect, much-branched
Leaves generally simple, alternate, sessile; blade linear to ovate, hairy, sometimes glandular, tip abruptly pointed
Inflorescence: flowers generally solitary in axils; pedicel bracts 2
Flower bisexual, bilateral; sepals 4–5, free, conspicuous; petals generally 5, 3 upper linear to clawed, held in ± upright "flag", 2 modified into glands flanking ovary; stamens generally 4, opening by pores; ovary superior, hairy, style slender, recurved
Fruit nut-like, bearing smooth or barbed spines
Genera in family: 1 genus, 17 species: Am, especially tropical
Reference: [Simpson 1989 Fl Neotropica 49:1–109]
Pollinating bees collect oils secreted by glandular petals.