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Jepson Interchange (more information)
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Richard R. Halse, except as specified; Robert W. Patterson, Family Editor

Annual, perennial herb, shrub, generally hairy, generally taprooted
Stem prostrate to erect
Leaves simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0
Inflorescence: cyme (generally raceme-like and coiled) or flowers solitary
Flower bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla generally deciduous, rotate to cylindric, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chamber 1, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2–5-chambered, styles 1–2, stigmas generally head-like
Fruit: capsule, generally loculicidal; valves generally 2
Genera in family: 20 genera, 300 species: especially w US; some cultivated (Emmenanthe, Nemophila, Phacelia )
Recent taxonomic note: Recently treated to be included in an expanded Boraginaceae (also including Lennoaceae) [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2000 Mol Phylog Evol 16:96–112]


Species in genus: 1 sp
Etymology: (Greek: 3 hearts, from calyx)


T. watsonii S. Watson


Perennial; herbage long-soft-hairy, becoming ± glabrous; taproot woody, generally topped by a branched caudex covered by persistent petiole bases of previous years
Stems several from root crown, erect, 5–40 cm
Leaves simple, generally in basal rosette, petioled, 2–9 cm, 5–25 mm wide; cauline alternate, smaller upward, lower short-petioled, upper sessile, 6–20 mm, 3–15 mm wide, entire
Inflorescence loose, terminal; flowers pedicelled
Flower: calyx lobes 5, very unequal, outer 3 cordate, 5–9 mm in flower, in fruit 9–25 mm, wide, scarious, veiny, green to purplish, inner 2 ± 4 mm in flower, linear, enlarged in fruit; corolla 4–8 mm, bell-shaped to rotate, white to cream, generally marked lavender; stamens included, unequal, equally attached; ovary chamber 1, style 1, 3–4 mm, tip lobed
Fruit 7–9 mm, glabrous
Seeds 4–8, oblong, brown, rough
Chromosomes: n=8
Ecology: Sandy or gravelly desert slopes, flats, mtns, generally in shelter of shrubs
Elevation: 100–2300 m.
Bioregional distribution: San Bernardino Mountains, East of Sierra Nevada, Desert
Distribution outside California: to Nevada, Utah, Arizona
Flowering time: Apr–Jun
Horticultural information: TRY.

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