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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Richard R. Halse, except as specified; Robert W. Patterson, Family Editor

Annual, perennial herb, shrub, generally hairy, generally taprooted
Stem prostrate to erect
Leaves simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0
Inflorescence: cyme (generally raceme-like and coiled) or flowers solitary
Flower bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla generally deciduous, rotate to cylindric, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chamber 1, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2–5-chambered, styles 1–2, stigmas generally head-like
Fruit: capsule, generally loculicidal; valves generally 2
Genera in family: 20 genera, 300 species: especially w US; some cultivated (Emmenanthe, Nemophila, Phacelia )
Recent taxonomic note: Recently treated to be included in an expanded Boraginaceae (also including Lennoaceae) [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2000 Mol Phylog Evol 16:96–112]


Annual, perennial herb, ± scapose; herbage ± glabrous to generally sparsely soft-hairy
Stem erect
Leaves simple; basal long-petioled, blades reniform to round, shallowly lobed or toothed; cauline few, reduced, alternate
Inflorescence loose; flowers on pedicels
Flower: corolla > calyx, bell- to funnel-shaped, white, generally yellow in center; stamens included, subequal; ovary chambers 2, style 1, thread-like, included, generally slightly lobed
Fruit oblong to ovoid
Seeds many, ovoid, angular, brown, pitted
Species in genus: 5 species: CA, AK, MT, w Can
Etymology: (Count N. von Romanzoff, promoter of Russian expedition to CA in 1816)


R. tracyi Jeps.

Plant 2–12 cm, forming rounded tufts; tubers clustered, ovoid, brown, tomentose
Leaf: petioles 1–8 cm, soft-hairy; base widened; blades 10–35 mm wide, ± glabrous to soft-hairy
Inflorescence compact, generally < leaves; pedicels stout, < 1 cm
Flower: calyx lobes 2–5 mm, lanceolate, acute, soft-hairy; corolla 6–8 mm; ovary soft-hairy, style 2–3 mm
Fruit 5–8 mm
Chromosomes: n=11
Ecology: Uncommon. Rocky ocean bluffs
Elevation: < 30 m.
Bioregional distribution: North Coast
Distribution outside California: to Washington
See the CNPS Inventory for information about endangerment and rarity.
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