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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

HYDROPHYLLACEAE

WATERLEAF FAMILY

Richard R. Halse, except as specified; Robert W. Patterson, Family Editor

Annual, perennial herb, shrub, generally hairy, generally taprooted
Stem prostrate to erect
Leaves simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0
Inflorescence: cyme (generally raceme-like and coiled) or flowers solitary
Flower bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla generally deciduous, rotate to cylindric, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chamber 1, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2–5-chambered, styles 1–2, stigmas generally head-like
Fruit: capsule, generally loculicidal; valves generally 2
Genera in family: 20 genera, 300 species: especially w US; some cultivated (Emmenanthe, Nemophila, Phacelia )
Recent taxonomic note: Recently treated to be included in an expanded Boraginaceae (also including Lennoaceae) [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2000 Mol Phylog Evol 16:96–112]

ROMANZOFFIA

Annual, perennial herb, ± scapose; herbage ± glabrous to generally sparsely soft-hairy
Stem erect
Leaves simple; basal long-petioled, blades reniform to round, shallowly lobed or toothed; cauline few, reduced, alternate
Inflorescence loose; flowers on pedicels
Flower: corolla > calyx, bell- to funnel-shaped, white, generally yellow in center; stamens included, subequal; ovary chambers 2, style 1, thread-like, included, generally slightly lobed
Fruit oblong to ovoid
Seeds many, ovoid, angular, brown, pitted
Species in genus: 5 species: CA, AK, MT, w Can
Etymology: (Count N. von Romanzoff, promoter of Russian expedition to CA in 1816)

Native

R. californica Greene

Plant 10–40 cm; tubers clustered, ovoid, brown, tomentose
Leaf: petioles 2–12 cm; base slightly widened; blade 8–45 mm wide
Inflorescence > leaves, sometimes glandular; pedicels 1–3 cm in fruit, slender
Flower: calyx lobes 2–5 mm, linear-lanceolate, generally acute, glabrous, sometimes glandular; corolla 5–12 mm; ovary sparsely soft-hairy, glandular; style 4–7 mm
Fruit 6–10 mm
Chromosomes: n=11
Ecology: Ocean bluffs, roadbanks, wet cliffs, moist rocky areas
Elevation: < 800 m.
Bioregional distribution: Northwestern California, n Central Coast, San Francisco Bay Area
Distribution outside California: to Washington
Synonyms: R. suksdorfii Greene misapplied
Horticultural information: DRN, summerDRY: 4, 5, 6, 17 &SHD: 7, 14, 15, 16, 19, 20, 21, 22, 23, 24.

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