This page is based on the 1993 Jepson Manual.
Please see the Jepson eFlora for up-to-date information about California vascular plants.
|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL||
previous taxon |
Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
Print edition is available from the University of California Press
|The second edition of The Jepson Manual (2012) is available from the University of California Press|
|See also the Jepson eFlora, which parallels the Second Edition|
Annual, perennial herb, shrub, generally hairy, generally taprooted
Stem prostrate to erect
Leaves simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0
Inflorescence: cyme (generally raceme-like and coiled) or flowers solitary
Flower bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla generally deciduous, rotate to cylindric, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chamber 1, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 25-chambered, styles 12, stigmas generally head-like
Fruit: capsule, generally loculicidal; valves generally 2
Genera in family: 20 genera, 300 species: especially w US; some cultivated (Emmenanthe, Nemophila, Phacelia )
Recent taxonomic note: Recently treated to be included in an expanded Boraginaceae (also including Lennoaceae) [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531553; Olmstead et al. 2000 Mol Phylog Evol 16:96112]
Annual, perennial herb, generally glandular-hairy, tap-rooted or from ± thick caudex
Leaves generally alternate, simple to 2-pinnately compound, generally ± reduced upward
Inflorescence: cyme, generally dense, coiled, generally 1-sided; pedicels generally short
Flower: corolla rotate to bell-shaped, white to purple, tube base with scales free or fused to filaments; stamens generally attached at same level, equal; ovary chamber 1 (or 2 below middle), placentas parietal, enlarging and meeting in fruit, style 2-lobed, generally hairy below lobes
Fruit oblong to spheric
Seeds 1many, oblong to spheric, generally brownish; back generally pitted or cross-furrowed
Species in genus: ± 175 species: Am; some cultivated for ornamental
Etymology: (Greek: cluster, from the dense inflorescence)
Reference: [Halse 1981 Madroño 28:121132; Heckard 1960 Univ Calif Publ Botany 32:1126; Lee 1988 Syst Bot 13:1620]
Bristly hairs may cause severe dermatitis. CA pers often hybridize, difficult to separate. Biennial and perennial herb species by Richard Halse.
Perennial 30150 cm
Stem prostrate to ascending, many-branched, glabrous to densely hairy, glandular or not
Leaf: blade 40200 mm, generally > petiole, oblong to widely ovate, compound; leaflets ± sessile, elliptic to oblong, coarsely toothed or lobed, lobes often toothed
Flower ± sessile; calyx lobes 46 mm, not generally longer in fruit, oblanceolate to ± spoon-shaped; corolla 58 mm, funnel- to bell-shaped, white to lavender, scales ovate; stamens 710 mm, exserted, glabrous; style 710 mm, exserted
Fruit 34 mm, ovoid, sharply bristly
Seeds 24, 23 mm, pitted
Ecology: Many habitats
Elevation: < 3100 m.
Bioregional distribution: Northwestern California (except Outer North Coast Ranges), High Cascade Range, Sierra Nevada, Central Western California, Southwestern California (except s Channel Islands), Warner Mountains, East of Sierra Nevada
Distribution outside California: to Washington, Idaho, Arizona
Varieties difficult, need study.
Stem generally < 60 cm, not glandular below inflorescence, densely puberulent; some hairs fine, long
Ecology: Slopes, meadows, coniferous forest
Elevation: 2003100 m.
Bioregional distribution: High Sierra Nevada, Tehachapi Mountain Area, San Gabriel Mountains, East of Sierra Nevada
Distribution outside California: to Oregon, Idaho
|YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).|