TREATMENT FROM THE JEPSON MANUAL	previous taxon | next taxon Jepson Interchange (more information)
	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

HYDROPHYLLACEAE

WATERLEAF FAMILY

Richard R. Halse, except as specified; Robert W. Patterson, Family Editor

Annual, perennial herb, shrub, generally hairy, generally taprooted
Stem prostrate to erect
Leaves simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0
Inflorescence: cyme (generally raceme-like and coiled) or flowers solitary
Flower bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla generally deciduous, rotate to cylindric, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chamber 1, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2–5-chambered, styles 1–2, stigmas generally head-like
Fruit: capsule, generally loculicidal; valves generally 2
Genera in family: 20 genera, 300 species: especially w US; some cultivated (Emmenanthe, Nemophila, Phacelia )
Recent taxonomic note: Recently treated to be included in an expanded Boraginaceae (also including Lennoaceae) [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2000 Mol Phylog Evol 16:96–112]

HESPEROCHIRON

Perennial, scapose; root caudex-like
Leaves simple, in basal rosette, spreading or ascending; blade tapered to petiole, generally entire, margins generally ciliate
Inflorescence: flower solitary; peduncle erect or spreading, 1–10 cm, slender
Flower: calyx lobes generally unequal, 2–9 mm, glabrous to hairy, ciliate; corolla tube generally densely hairy inside, throat generally yellow, lobes glabrous to hairy, white or bluish, generally tinged or marked with lavender or purple; stamens included, generally unequal, filament base widened; ovary hairy, chamber 1, style 1, stigmas 2, 2–5 mm
Fruit 5–11 mm, ovoid, hairy
Seeds many, ovoid, angular, reddish brown; surface honeycombed or pitted
Species in genus: 2 species: w US, n Mex
Etymology: (Greek: evening or western centaur)

Native

H. californicus (Benth.) S. Watson

Leaves generally > 6, < 8 cm, < 3 cm wide, oblanceolate to elliptic or ovate; surfaces generally densely to sparsely spreading-hairy
Inflorescence: flowers generally > 5
Flower: corolla < 3 cm, < 2 cm wide, bell- or funnel-shaped, lobes oblong, 3–10 mm
Chromosomes: n=8
Ecology: Wet meadows, flats, valleys
Elevation: 1000–2900 m.
Bioregional distribution: Klamath Ranges, High Cascade Range, High Sierra Nevada, Tehachapi Mountain Area, Western Transverse Ranges, San Bernardino Mountains, Great Basin Floristic Province
Distribution outside California: to Washington, Montana, Wyoming, Utah, Baja California
Flowering time: May–Jul
Horticultural information: IRR or WET, SUN, DRN: 1, 2, 3, 7, 18; DFCLT.