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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

HYDROPHYLLACEAE

WATERLEAF FAMILY

Richard R. Halse, except as specified; Robert W. Patterson, Family Editor

Annual, perennial herb, shrub, generally hairy, generally taprooted
Stem prostrate to erect
Leaves simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0
Inflorescence: cyme (generally raceme-like and coiled) or flowers solitary
Flower bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla generally deciduous, rotate to cylindric, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chamber 1, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2–5-chambered, styles 1–2, stigmas generally head-like
Fruit: capsule, generally loculicidal; valves generally 2
Genera in family: 20 genera, 300 species: especially w US; some cultivated (Emmenanthe, Nemophila, Phacelia )
Recent taxonomic note: Recently treated to be included in an expanded Boraginaceae (also including Lennoaceae) [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2000 Mol Phylog Evol 16:96–112]

EUCRYPTA

Annual, glandular, sticky, scented
Stem erect, much-branched
Leaves simple, 1–3-pinnately toothed to dissected; lower cauline leaves opposite, petioled; upper leaves alternate, becoming smaller, sessile, clasping; petioles generally narrowly winged, ciliate
Inflorescence terminal or axillary; pedicels thread-like, elongate in fruit
Flower: calyx < half-fused, bell-shaped, glandular, lobes oblong to spoon-shaped, ciliate; corolla bell-shaped, generally > calyx, with V-shaped transverse fold between each pair of filaments below throat; stamens included, equal, equally attached; ovary chamber 1 (or appearing 5 from complex, enlarged placenta), ovules borne on both sides of placenta, style 1, stigmas 2
Fruit ovoid to spheric, bristly
Seeds 5–15
Species in genus: 2 species: sw US
Etymology: (Greek: well hidden, from seeds)
Reference: [Constance 1938 Lloydia 1:143–152]

Native

E. chrysanthemifolia (Benth.) Greene

Stem erect to openly spreading, < 9 dm
Leaves: lower 2–10 cm, 1–5 cm wide, petioles < 1/2 blade, widened, clasping, blade oblong to widely ovate, pinnate to deeply pinnately lobed, lobes 7–13, deeply 1–2-pinnately lobed, teeth obtuse; upper leaves smaller, narrower, less lobed, bases clasping
Inflorescence: flowers 4–15 per branch; pedicels generally recurved in fruit
Flower: calyx 2–4 mm; corolla 2–6 mm, lobes hairy on back; style < 3 mm
Fruit 2–4 mm wide, < spreading calyx
Seeds 6–8, dark brown, of 2 kinds; some elliptic or round, disk-like, smooth; others oblong-ovoid, wrinkled
Ecology: Canyons, chaparral, disturbed areas, slopes
Elevation: 0–2300 m.
Bioregional distribution: s Sierra Nevada Foothills, Tehachapi Mountain Area, San Joaquin Valley, Central Western California, Southwestern California, East of Sierra Nevada, Desert
Distribution outside California: Nevada, Arizona, Baja California

Native

var. chrysanthemifolia

Stem erect, stout
Leaves: lower 2–10 cm, 1–5 cm wide, lobes 9–13
Inflorescence: flowers 8–15 per branch
Flower: corolla 3–6 mm, 4–8 mm wide, > calyx, yellowish white; style 1–3 mm
Chromosomes: n=10
Ecology: Roadsides, burns, coastal bluffs, ravines
Elevation: 0–1000 m.
Bioregional distribution: Tehachapi Mountain Area, s San Joaquin Valley, Central Western California, Southwestern California
Distribution outside California: Baja California