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EUPHORBIACEAE

SPURGE FAMILY

Grady L. Webster, except as specified

Annual, perennial herb, shrub, tree, vine, monoecious or dioecious
Stem generally branched, sometimes fleshy or spiny
Leaves generally simple, alternate or opposite, generally stipuled, petioled; blade entire, toothed, or palmately lobed
Inflorescence: cyme, panicle, raceme, spike; flowers sometimes in clusters (dense, enclosed by involucre, flower-like in Chamaesyce, Euphorbia ), terminal or axillary
Flower unisexual, ± radial; sepals generally 3–5, free or fused; petals generally 0; stamens 1–many, free or filaments fused; ovary superior, chambers 1–4, styles free or fused, simple or lobed
Fruit: generally capsule
Seeds 1–2 per chamber; seed scar appendage sometimes present, pad- to dome-like
Genera in family: 300 genera, 7500 species: ± worldwide especially tropical; some cultivated (Aleurites , tung oil; Euphorbia subsp.; Hevea , rubber; Ricinus )
Reference: [Webster 1967 J Arnold Arbor 48:303–430]
Many species ± highly TOXIC.

DITAXIS

Annual, perennial herb, subshrub, generally monoecious; sap clear; hairs 0 or generally 2-forked, generally appressed
Stems spreading to erect, 1–10 dm
Leaves simple, alternate, stipuled
Inflorescence: raceme, axillary; staminate flowers generally above pistillate flowers; axis generally densely appressed-hairy; bracts entire
Staminate flower: sepals 5, edges abutting in bud; petals 5; stamens 5–15, generally in 2 sets, some > others, filaments fused into a column, staminodes 0–3 at column tip
Pistillate flower: sepals 5, overlapping in bud; petals 5; nectar disk ± dissected; ovary 3-chambered, styles 3, 2-lobed
Fruit smooth
Seeds: surface net-like to finely pitted; scar not appendaged
Species in genus: ± 50 species: tropical, warm temp Am
Etymology: (Greek: 2-ranked, from 2 sets of anthers)

Native

D. lanceolata (Benth.) Pax & K. Hoffm.

Subshrub
Stems generally erect, 1–5 dm, brittle, appressed-hairy
Leaf 2–6 cm; stipules ± 1 mm, entire; blade lanceolate, densely hairy, margin entire
Staminate flower: sepals 2.5–3 mm, hairy; petals 3–3.5 mm, back hairy; stamen column ± 1.5 mm
Pistillate flower: sepals 3–4 mm, entire; petals ± = sepals, lanceolate to ovate, back hairy; ovary densely appressed-hairy, styles generally free, lobe tips expanded
Fruit 3–5 mm
Seed 2–2.5 mm, angled, pitted
Ecology: Rocky soils, slopes, canyons
Elevation: < 600 m.
Bioregional distribution: Mojave Desert (Eagle Mtn), Sonoran Desert
Distribution outside California: Arizona, Mexico
Flowering time: Mar–May

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