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EUPHORBIACEAE

SPURGE FAMILY

Grady L. Webster, except as specified

Annual, perennial herb, shrub, tree, vine, monoecious or dioecious
Stem generally branched, sometimes fleshy or spiny
Leaves generally simple, alternate or opposite, generally stipuled, petioled; blade entire, toothed, or palmately lobed
Inflorescence: cyme, panicle, raceme, spike; flowers sometimes in clusters (dense, enclosed by involucre, flower-like in Chamaesyce, Euphorbia ), terminal or axillary
Flower unisexual, ± radial; sepals generally 3–5, free or fused; petals generally 0; stamens 1–many, free or filaments fused; ovary superior, chambers 1–4, styles free or fused, simple or lobed
Fruit: generally capsule
Seeds 1–2 per chamber; seed scar appendage sometimes present, pad- to dome-like
Genera in family: 300 genera, 7500 species: ± worldwide especially tropical; some cultivated (Aleurites , tung oil; Euphorbia subsp.; Hevea , rubber; Ricinus )
Reference: [Webster 1967 J Arnold Arbor 48:303–430]
Many species ± highly TOXIC.

BERNARDIA

Shrub, monoecious or dioecious; sap clear
Stems erect, generally much-branched
Leaves simple, cauline, alternate, stipuled; hairs simple or stellate
Staminate inflorescence: spike or raceme, axillary
Pistillate inflorescence terminal; flower sometimes solitary
Staminate flower sessile or short-pedicelled; calyx splitting into 3–4 segments; petals 0; stamens 3–25, filaments free; nectar disk minute or 0
Pistillate flower sessile; sepals 4–6; petals 0; nectar disk 0; ovary 3-chambered, styles 3, free, 2-lobed or -toothed
Fruit 3-lobed
Seed 1 per chamber; scar appendaged
Species in genus: 30–40 species: tropical, subtropical Am
Etymology: (Latin: Bernard de Jussieu, French taxonomist, 1699–1776)

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