|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL (1993)||
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY
Annual, biennial, perennial herb (rarely shrub, tree), often from taproot
Stem often ± scapose, generally ribbed, hollow
Leaves basal and generally some cauline, generally alternate; stipules generally 0; petiole base generally sheathing stem; blade generally much dissected, sometimes compound
Inflorescence: umbel or head, simple or compound, generally peduncled; bracts present (in involucres) or not; bractlets generally present (in involucels)
Flowers many, small, generally bisexual (or some staminate), generally radial (or outer bilateral); calyx 0 or lobes 5, small, atop ovary; petals 5, free, generally ovate or spoon-shaped, generally incurved at tips, generally ± ephemeral; stamens 5; pistil 1, ovary inferior, 2-chambered, generally with a ± conic, persistent projection or platform on top subtending 2 free styles
Fruit: 2 dry, 1-seeded halves that separate from each other but generally remain attached for some time to a central axis; ribs on each half 5, 2 marginal and 3 on back; oil tubes 1several per interval between ribs
Genera in family: 300 genera, 3,000 species: ± worldwide, especially temp; many cultivated for food or spice (e.g., Carum, caraway; Daucus; Petroselinum); some highly toxic (e.g., Conium). Underground structures here called roots, but true nature remains problematic. Mature fruit generally critical in identification; shapes generally given in outline, followed by shape in X -section of 2 fruit halves together.
Perennial, taprooted, generally glabrous
Stem generally 0 or short
Leaves mostly basal, membranous to subleathery or fleshy; blade oblong to widely ovate or round, palmately or pinnately lobed to 12-pinnately or -ternate-pinnately dissected or compound, segments or leaflets linear to obovate, entire to variously lobed, generally spine-tipped
Inflorescence: umbels compound, generally terminal, scapose, open to spheric, dense, peduncled; bracts, bractlets conspicuous and scarious (or rarely 0); rays fewmany (rays and pedicels sometimes ± 0)
Flower: calyx lobes prominent to 0; petals oblong to obovate, white, yellow, or purple, tips narrowed; projection atop ovary 0
Fruit oblong to ovate, subcylindric to compressed front-to-back; ribs subequal or unequal, marginal and some or all others thin- or corky-winged, or rarely some or all wingless; oil tubes per rib-interval 1several; fruit axis 0 or divided to base
Seed: face flat to longitudinally concave or grooved
Species in genus: ± 50 species: w North America
Etymology: (Greek: wave wing)
Reference: [Mathias 1930 Ann Missouri Bot Gard 17:213476]
Generic boundaries fluctuating.
Some species outside CA are TOXIC to livestock.
See the CNPS Inventory for information about endangerment and rarity.
Stem 1223 cm, glabrous, glaucous; base fibrous
Leaf: petiole 818 cm; blade 2.54.5 cm, round-reniform, ternate, leaflets obovate, deeply lobed, ultimate lobes toothed, spine-tipped
Inflorescence: peduncles > leaves; bracts 0; bractlets inconspicuous, leaf-like, linear to lanceolate; rays ± 8, 12 cm; pedicels 25 mm
Flower: corolla purplish
Fruit 78 mm, widely ovate; ribs unequal, marginal and 1 or 2 other wings > body in width
Ecology: Limestone, gypsum slopes
Elevation: 10002000 m.
Bioregional distribution: ne Desert Mountains
Distribution outside California: Nevada
Flowering time: AprMay