Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



James Henrickson

Annual to tree
Stem prostrate to erect
Leaves alternate or opposite, simple, generally entire, bract-like upward; stipules 0
Inflorescence: cymes, often arrayed ± in spikes or panicles; bracts 1–3 per flower, generally membranous-scarious, tip often short-pointed or spine-like
Flower generally small, uni- or bisexual, radial; sepals 0–5, fused at base or free, often ± scarious; petals 0; stamens 0–5, opposite sepals (staminodes sometimes alternating), filaments sometimes fused at base; ovary superior, 1-chambered, ovules 1–several, erect or pendent on ± basal stalks, styles 0–3, stigma head-like or 2–3-lobed
Fruit: utricle to circumscissile capsule
Seed lenticular to spheric, hard
Genera in family: ± 65 genera, ± 900 species: tropical, subtropical
Reference: [Robertson 1981 J Arnold Arbor 62:267–313]



Annual, monoecious or dioecious
Leaves alternate; blade linear to ovate
Inflorescence: cymes in dense, spike-like clusters; bract 1, tip generally short-pointed or -spined, generally scarious; bractlets 0–2
Staminate flower: sepals 3–5, ± equal, ± as bracts; stamens (1)3–5, filaments free; staminodia 0
Pistillate flower: sepals (3–)5, ± equal, scarious except midvein, fused at base, falling with fruit; ovary compressed-ovoid, styles (2)3, stigmas slender, papillate, ovule 1, erect
Fruit circumscissile or indehiscent, smooth or inflated-wrinkled; walls membranous to spongy-hardened
Seed 1, lenticular, smooth, reddish to black
Species in genus: ± 60 species: worldwide; some potherbs, some cultivated for seed
Etymology: (Greek: unfading, from persistent bracts and sepals)
Some species (especially A. cruentus, A. powellii, A. retroflexus) hybridize complexly
Reference: [Tucker & Sauer Madroño 1958 14:252–261]


A. powellii S. Watson

Plant erect, 3–20 dm, monoecious
Leaf: petiole 10–50 mm; blade 15–90 mm, lower diamond-shaped to ovate, upper ± lanceolate
Inflorescence: panicle of spike-like clusters 1–1.8 cm wide; axillary clusters 1–6 cm; terminal clusters < or = 25 cm; bracts 3–6(8) mm, spine-tipped, midrib green, thick, margin scarious in lower 2/3
Staminate flower: sepals 4–5, 2–3 mm; stamens 3–5
Pistillate flower: sepals 5, 1.7–3.6 mm, unequal, ± oblong, tip short-pointed
Fruit 2.5–3.6 mm, circumscissile, smooth throughout
Seed 0.8–1.4 mm wide, reddish black
Ecology: Uncommon. Waste places
Elevation: < 800 m.
Bioregional distribution: Sierra Nevada Foothills, Great Central Valley, Central Western California, Southwestern California, White and Inyo Mountains
Distribution outside California: to Washington, north-central US, Texas, Mexico, w S.America
Flowering time: Jul–Oct

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