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CAMPANULACEAE

BELLFLOWER FAMILY

Nancy Morin, except as specified

Annual to tree
Leaves generally cauline, generally simple, generally alternate, petioled or not; stipules 0
Inflorescence: panicle, raceme, spike, or flowers solitary in axils, generally open; bracts leaf-like or not
Flower: bisexual, radial or bilateral, sometimes inverted (pedicel twisted 180°; hypanthium generally present, ± fused to ovary; sepals generally 5; corolla radial to 2-lipped, generally fused (tube sometimes split down back), lobes generally 5; stamens 5, free or ± fused (anthers and filaments fused into tube or filaments fused above middle); ovary inferior, sometimes half inferior, chambers 1–3, placentas axile or parietal, ovules many, style generally 1, 2–5-branched
Fruit: generally capsule, dehiscing on sides or at tip by pores or short valves
Seeds many
Genera in family: ± 70 genera, ± 2000 species: worldwide. Some cultivated for ornamental (Campanula, Jasione, Lobelia ). Subfamilies sometimes treated as different families.

NEMACLADUS

Nancy R. Morin and Jennifer Milburn

Annual; roots fibrous
Stems erect or spreading, simple or branched at base or below middle
Leaves basal; petiole short or 0
Inflorescence ± raceme-like; bract 1 per flower, small; pedicel generally thread-like
Flower inverted; sepals linear to triangular; corolla nearly radial and 5-lobed or 2-lipped (upper lip 3-lobed, lower lip 2-lobed); filaments free at base, fused into tube above, sometimes appendaged at tube base, anthers free, all alike; ovary generally half-inferior in fruit, sometimes 0, hemispheric to obconic, sometimes glandular, stigma 2-lobed, papillate
Fruit generally > hypanthium, hemispheric to fusiform; tip pointed or rounded, dehiscing at tip by 2 valves; chambers 2
Seed elliptic to oblong
Species in genus: 13 species: sw US, nw Mex
Etymology: (Greek: thread-like branch)
Reference: [McVaugh 1942 N Amer Flora 32A: 1–134]

Native

N. capillaris Greene


Stems stiffly ascending, 7–18 cm; base brownish or purplish
Leaf 3–15 mm, ovate, narrowed abruptly to short petiole, entire, glabrous or hairy
Inflorescence: axis zigzag, especially in fruit; bracts 1–3 mm, lanceolate to elliptic; pedicels 8–12 mm, 0.1–0.15 mm diam, spreading, straight or slightly curved, tip not curved
Flower: hypanthium ± 1 mm; sepals ± 0.5 mm, elliptic to lanceolate, erect; corolla 0.7–1.3 mm, divided ± to base, white, glabrous; filament tube ± 1 mm, tip slightly curved, glabrous, anthers 0.1–0.2 mm
Fruit 1.5–2.5 mm, obconic (base acute, tip rounded)
Seed 0.5–0.7 mm, widely elliptic, narrowly ridged
Chromosomes: n=9
Ecology: Dry slopes, burned areas
Elevation: 400–2100 m.
Bioregional distribution: North Coast Ranges, High Cascade Range, n Sierra Nevada Foothills, High Sierra Nevada, Modoc Plateau, Mojave Desert
Distribution outside California: Oregon
Flowering time: May–Jul

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