TREATMENT FROM THE JEPSON MANUAL	previous taxon | next taxon Jepson Interchange (more information)
	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

BRASSICACEAE

MUSTARD FAMILY

Reed C. Rollins, except as specified

Annual to subshrub
Leaves generally basal and cauline, alternate, generally simple; stipules 0
Inflorescence: generally raceme
Flower bisexual; sepals 4, free; petals (0)4, free, generally white or yellow, often clawed; stamens generally (2,4)6, generally 4 long, 2 short; ovary 1, superior, chambers generally 2, septum membranous, connecting 2 parietal placentas, style 1, stigma simple or 2-lobed
Fruit: generally capsule ("silique") with 2 deciduous valves, sometimes breaking transversely or indehiscent
Seeds 1–many per chamber
Genera in family: 300+ genera, 3000+ species: worldwide, especially cool regions; some cultivated for food (especially Brassica, Raphanus ) and ornamental
Recent taxonomic note: Recently treated to include Capparaceae [Rodman et al. 1993 Ann Missouri Bot Gard 80:686–699; Rollins 1993 Cruciferae of Continental North America. Stanford Univ Press]
Family description, key to genera by Robert A. Price.

PHYSARIA

DOUBLE BLADDERPOD

Perennial, silvery; taproot enlarged; hairs stellate, generally ± sessile, appressed
Stems several–many, generally from below rosette; outer prostrate to decumbent
Leaves: basal obovate to ± round, < petioles; cauline entire, upper generally sessile
Flower: petals yellow
Fruit widely obovate to ± round, inflated; tip notched to ± middle, generally also notched from base; valves shed with seeds enclosed; style persistent
Seeds 2–6 per chamber, ± flat; margins 0; embryonic root at edges of both cotyledons
Species in genus: 22 species: w North America, generally Rocky Mtns, Great Basin
Etymology: (Greek: bellows, from inflated fruit)

Native

P. chambersii Rollins

Plant cespitose
Stem unbranched, 5–15 cm
Leaves: basal 3–6 cm, 1–2 cm wide, entire to dentate, tip obtuse; cauline 1–2 cm, spoon-shaped, tip generally acute
Inflorescence 2–10 cm, dense, ± umbel-like
Fruit 1–1.5 cm, hairy; notch in base shallow or ± 0; septum 4–6 mm; pedicel ascending to spreading, 8–15 mm; style 6–8 mm, exserted from notch
Seeds generally 4 per chamber
Chromosomes: 2n=8, 10,16,24
Ecology: Limestone soils
Elevation: 1500–2500 m.
Bioregional distribution: n Desert Mountains (Clark, Grapevine mtns)
Distribution outside California: to Oregon, Utah, Arizona
Flowering time: May
Horticultural information: TRY; DFCLT.

See the CNPS Inventory for information about endangerment and rarity.
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Retrieve Jepson Interchange Index to Plant Names entry for Physaria chambersii
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