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This page is based on the 1993 Jepson Manual.
Please see the Jepson eFlora for up-to-date information about California vascular plants. |
| Jepson Flora Project: Jepson Interchange |
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TREATMENT FROM THE JEPSON MANUAL |
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Jepson Interchange (more information) |
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©Copyright 1993 by the Regents of the University of California
Print edition is available from the University of California Press |
| The second edition of The Jepson Manual (2012) is available from the University of California Press | |
| See also the Jepson eFlora, which parallels the Second Edition |
Annual, perennial herb, shrubs, generally bristly or sharply hairy
Stem prostrate to erect
Leaves cauline, often with basal rosette, generally simple, alternate; lower sometimes opposite, entire
Inflorescence: cyme, generally elongate, panicle-, raceme- or spike-like, coiled in flower, generally uncoiled in fruit or flowers 12 per axil
Flowers generally bisexual, generally radial; sepals 5, free or fused in lower half; corolla 5-lobed, generally salverform, top of tube generally appendaged, appendages 5, alternating with stamens, sometimes arching over tube; stamens 5, epipetalous; ovary superior, generally 4-lobed, style generally entire
Fruit: nutlets 14, smooth to variously roughened, sometimes prickly or bristled
Genera in family: ± 100 genera, ± 2000 species: tropical, temp, especially w North America, Medit; some cultivated (Borago, Echium, Myosotis, Symphytum ).Almost all genera may be TOXIC from alkaloids or accumulated nitrates
Recent taxonomic note: Recently treated to include Hydrophyllaceae [Olmsted et al. 2000 Mol Phylog Evol 16:96112]
Family description, key to genera by Timothy C. Messick.
Annual, perennial herb, hairy, taprooted
Stem erect
Leaves generally cauline, ± sessile, entire
Inflorescence: bracted cymes, open panicles, or flowers solitary in upper leaf axils
Flower: calyx deeply 5-lobed, enlarging in fruit; corolla 5-lobed, funnel-shaped or salverform, generally ± yellow, appendages present or not; style entire
Fruit: nutlets 14, 36 mm, smooth to pitted or wrinkled
Species in genus: 75 species: worldwide, generally temp or mtn
Etymology: (Greek: stone seed)
Reference: [Baker 1961 Rhodora 63:229235]
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Perennial, strigose; caudex woody
Stems fewseveral, 13 dm, clustered, ± unbranched
Leaves many; blade 1.56 cm, linear to linear-oblong
Inflorescence: cymes many, in upper axils; pedicels 25 mm, ± recurved in fruit
Flower: corolla 1535 mm, 23.5 X calyx, 1020 mm wide, salverform, yellow, appendaged
Fruit: nutlets ± pitted, shiny, gray
Chromosomes: 2n=24,36
Ecology: Sandy, rocky slopes, pinyon/juniper woodland
Elevation: 16501700 m.
Bioregional distribution: Desert Mountains (Keystone Canyon, New York Mtns, San Bernardino Co.)
Distribution outside California: to British Columbia, Montana, Great Plains, s Nevada
Not heterostylous; cleistogamous flowers presentHorticultural information: DRN: 1, 2, 3, 16, 17; DFCLT.
| YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps). |
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