|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL (1993)||
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY
Annual, perennial herb, shrubs, generally bristly or sharply hairy
Stem prostrate to erect
Leaves cauline, often with basal rosette, generally simple, alternate; lower sometimes opposite, entire
Inflorescence: cyme, generally elongate, panicle-, raceme- or spike-like, coiled in flower, generally uncoiled in fruit or flowers 12 per axil
Flowers generally bisexual, generally radial; sepals 5, free or fused in lower half; corolla 5-lobed, generally salverform, top of tube generally appendaged, appendages 5, alternating with stamens, sometimes arching over tube; stamens 5, epipetalous; ovary superior, generally 4-lobed, style generally entire
Fruit: nutlets 14, smooth to variously roughened, sometimes prickly or bristled
Genera in family: ± 100 genera, ± 2000 species: tropical, temp, especially w North America, Medit; some cultivated (Borago, Echium, Myosotis, Symphytum ).
Almost all genera may be TOXIC from alkaloids or accumulated nitrates
Recent taxonomic note: Recently treated to include Hydrophyllaceae [Olmsted et al. 2000 Mol Phylog Evol 16:96112]
Family description, key to genera by Timothy C. Messick.
Annual, perennial herb, hairy, taprooted
Leaves generally cauline, ± sessile, entire
Inflorescence: bracted cymes, open panicles, or flowers solitary in upper leaf axils
Flower: calyx deeply 5-lobed, enlarging in fruit; corolla 5-lobed, funnel-shaped or salverform, generally ± yellow, appendages present or not; style entire
Fruit: nutlets 14, 36 mm, smooth to pitted or wrinkled
Species in genus: 75 species: worldwide, generally temp or mtn
Etymology: (Greek: stone seed)
Reference: [Baker 1961 Rhodora 63:229235]
Perennial, strigose; caudex woodySee the CNPS Inventory for information about endangerment and rarity.
Stems fewseveral, 13 dm, clustered, ± unbranched
Leaves many; blade 1.56 cm, linear to linear-oblong
Inflorescence: cymes many, in upper axils; pedicels 25 mm, ± recurved in fruit
Flower: corolla 1535 mm, 23.5 X calyx, 1020 mm wide, salverform, yellow, appendaged
Fruit: nutlets ± pitted, shiny, gray
Ecology: Sandy, rocky slopes, pinyon/juniper woodland
Elevation: 16501700 m.
Bioregional distribution: Desert Mountains (Keystone Canyon, New York Mtns, San Bernardino Co.)
Distribution outside California: to British Columbia, Montana, Great Plains, s Nevada
Not heterostylous; cleistogamous flowers present
Horticultural information: DRN: 1, 2, 3, 16, 17; DFCLT.