Jepson Flora Project: Jepson Interchange    

 
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
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BORAGINACEAE

BORAGE FAMILY

Annual, perennial herb, shrubs, generally bristly or sharply hairy
Stem prostrate to erect
Leaves cauline, often with basal rosette, generally simple, alternate; lower sometimes opposite, entire
Inflorescence: cyme, generally elongate, panicle-, raceme- or spike-like, coiled in flower, generally uncoiled in fruit or flowers 1–2 per axil
Flowers generally bisexual, generally radial; sepals 5, free or fused in lower half; corolla 5-lobed, generally salverform, top of tube generally appendaged, appendages 5, alternating with stamens, sometimes arching over tube; stamens 5, epipetalous; ovary superior, generally 4-lobed, style generally entire
Fruit: nutlets 1–4, smooth to variously roughened, sometimes prickly or bristled
Genera in family: ± 100 genera, ± 2000 species: tropical, temp, especially w North America, Medit; some cultivated (Borago, Echium, Myosotis, Symphytum ). Almost all genera may be TOXIC from alkaloids or accumulated nitrates
Recent taxonomic note: Recently treated to include Hydrophyllaceae [Olmsted et al. 2000 Mol Phylog Evol 16:96–112]
Family description, key to genera by Timothy C. Messick.

HACKELIA

STICKSEED

Robert L. Carr

Perennial (rarely biennial); hairs appressed to spreading; caudex generally branched in age, often ± woody, taprooted
Stem ascending or erect
Leaves: lowest generally with petioles ± = blades, ± winged; other leaves generally sessile, becoming bract-like toward inflorescence
Inflorescence: cymes, generally ± terminal, generally > 3, generally arrayed in panicles, coiled at tips
Flower radial; corolla rotate-salverform, with appendage near base of each lobe
Fruit: nutlets erect, > style, generally with lateral-medial scar, generally with barb-tipped prickles on margin and exposed face
Species in genus: 40 species: generally w North America, se Asia
Etymology: (J. Hackel, Czech botanist, born 1783)
Reference: [Gentry & Carr 1976 Mem New York Bot Gard 26:121–227]
Difficult genus needing much work, especially in n CA, se Asia; sometimes merged with Lappula
Horticultural information: TRY; DFCLT.

Native

H. californica (A. Gray) I.M. Johnst.


Stem 3–10 dm; hairs strigose to ± long, soft
Leaves: caudex, lowest cauline 3–17 cm, 6–30 mm wide, lanceolate to oblanceolate; mid to upper cauline lanceolate to ovate, often clasping
Flower: corolla white or rarely pinkish, limb 5–14 mm wide, appendages generally wider than long
Fruit: nutlets 4–4.5 mm, facial prickles 10–26, ± as long as marginal
Chromosomes: 2n=24
Ecology: Meadows, forest openings
Elevation: 900–1800 m.
Bioregional distribution: High North Coast Ranges, High Cascade Range, n High Sierra Nevada.

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bioregional map for HACKELIA%20californica being generated
 
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Retrieve Jepson Interchange Index to Plant Names entry for Hackelia californica
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