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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

BERBERIDACEAE

BARBERRY FAMILY

Michael P. Williams

Perennial, shrub, generally from rhizomes; caudex sometimes present, glabrous, glaucous, or hairy
Stems spreading to erect, branched or not
Leaves simple, 1–3-ternate, or pinnately compound, basal and cauline, generally alternate, deciduous or evergreen, petioled
Inflorescence: generally raceme, spike, or panicle, scapose, terminal, or axillary
Flower: sepals 6–18 or 0, generally in whorls of 3; petals generally 6, in 2 whorls of 3, or 0; stamens 6–12, free or fused at base, 2-whorled or not, anthers dehiscent by flap-like valves or longitudinal slits; ovary superior, chamber 1, ovules generally 1–10, style 1 or 0, stigma flat or spheric
Fruit: berry, capsule, or achene
Genera in family: 16 genera, ± 670 species: temp, tropical worldwide; some cultivated (Berberis, Epimedium, Nandina (Heavenly bamboo), Vancouveria )
Reference: [Ernst 1964 J Arnold Arbor 45:1–35]

ACHLYS

VANILLA LEAF, DEER FOOT

Perennial from scaly rhizomes; caudex short, erect
Leaves 1–few, basal, long-petioled, ternate; leaflets triangular to fan-shaped, base tapered
Inflorescence: spike, ± scapose, dense, long-peduncled; lateral flowers generally unisexual, terminal flowers bisexual
Flower: perianth 0; stamens generally 9, anther valves flap-like, curled inward; ovule 1, style 0, stigma ± flat, furrowed
Fruit: achene, curved, furrowed, brown to red-purple
Species in genus: 2 species: w North America, Japan
Etymology: (Greek: thin mist or obscurity, from inconspicuous flowers)
Reference: [Terabayashi 1981 Bot Mag Tokyo 94:141–157]

Native

A. triphylla (Sm.) DC. subsp. triphylla

Plant 2–4 dm; rhizome internodes 2.5–5 cm
Leaf: terminal leaflet generally 4–11 cm, 4–8 cm wide, 3(–5)-lobed
Inflorescence (except peduncle) 2.5–5 cm
Flower: stamens 3–4 mm; ovary 1–1.5 mm
Fruit 3–4.5 mm, red-purple
Chromosomes: 2n=12
Ecology: Moist shaded sites, coniferous forest
Elevation: < 1500 m.
Bioregional distribution: North Coast, w Klamath Ranges, w Outer North Coast Ranges
Distribution outside California: to British Columbia
Other subsp. in Japan
Horticultural information: SHD, IRR, DRN: 4, 5, 6, 15, 16, 17; GRCVR; DFCLT.

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