| TREATMENT FROM THE JEPSON MANUAL (1993) |
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©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora. |
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY |
Perennial, in soil or rock crevices; rhizome generally short-creeping, suberect, or erect, scales large, generally tan to brown, generally 1-colored
Leaves generally tufted, 5–200+ cm, generally ± alike; petiole generally firm, base generally darker, with 2–many vascular strands; blade 1–4-pinnate, often with scales, hair-like scales, hairs (except clear, needle-like hairs generally 0), or short-stalked glands on axes, sometimes between veins, veins free to netted; 1° and 2° axes generally grooved on upper side
Sporangia: sori round, less often oblong or J-shaped, along or at tips of veins; indusia peltate, round-reniform, oblong to linear, J-shaped, hood-like, or cup-like, rarely 0; spores elliptic, winged, ridged, or spiny, scar linear
Genera in family: ± 60 genera, > 1000 species: worldwide, especially tropical, wooded areas. Woodsia sometimes in Woodsiaceae; Athyrium, Cystopteris sometimes in Athyriaceae.
Rhizome short-creeping or ascending to suberect, stout
Leaf: petiole stout, base firm, scaly, in X -section with many ± round or ovate vascular strands in an arc; blade 1-pinnate, generally thick, leathery, veins regularly netted
Sporangia: sori round; indusium peltate, not persistent
Species in genus: ± 20 species: generally eastern Asia
Etymology: (Greek: arch, from pattern of netted veins)
| Introduced |
Rhizome with old, large, light- to dark-brown, entire to jagged scales
Leaf 30–80 cm; 1° leaflets 8–11 cm, often with 1 lobe on distal side of base, teeth 0–10 mm, < 6 mm wide, without bristle-like tips, margin thickened
Chromosomes: 2n=164
Ecology: Generally moist cliffs, banks, crevices
Elevation: < 900 m.
Bioregional distribution: Outer North Coast Ranges, n Sierra Nevada Foothills, San Joaquin Valley, Central Coast, Outer South Coast Ranges, South Coast, Western Transverse Ranges, San Gabriel Mountains, Peninsular Ranges (Santa Ana Mtns)
Distribution outside California: also se US; native to eastern Asia
cultivated as ornamental.