Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Alan R. Smith and Thomas Lemieux

Perennial, in soil or rock crevices; rhizome generally short-creeping, suberect, or erect, scales large, generally tan to brown, generally 1-colored
Leaves generally tufted, 5–200+ cm, generally ± alike; petiole generally firm, base generally darker, with 2–many vascular strands; blade 1–4-pinnate, often with scales, hair-like scales, hairs (except clear, needle-like hairs generally 0), or short-stalked glands on axes, sometimes between veins, veins free to netted; 1° and 2° axes generally grooved on upper side
Sporangia: sori round, less often oblong or J-shaped, along or at tips of veins; indusia peltate, round-reniform, oblong to linear, J-shaped, hood-like, or cup-like, rarely 0; spores elliptic, winged, ridged, or spiny, scar linear
Genera in family: ± 60 genera, > 1000 species: worldwide, especially tropical, wooded areas. Woodsia sometimes in Woodsiaceae; Athyrium, Cystopteris sometimes in Athyriaceae.



Rhizome short-creeping to suberect, stout
Leaf: petiole stout, fleshy, easily crushed, straw-colored except base generally blackened, base scaly, in X -section with 2 crescent-shaped vascular strands; blade generally 2-pinnate or more, ± glabrous, veins free
Sporangia: sori ± round, ± oblong, or J-shaped; indusium oblong or J-shaped, laterally attached, or 0
Species in genus: ± 100 species: generally n temp, especially e Asia
Etymology: (Greek: doorless, from enclosed sori)


A. alpestre (D.H. Hoppe) Clairv. var. americanum Butters

Leaf: blade narrow-elliptic or lanceolate, 2–3-pinnate, segments shallowly to deeply pinnately lobed, marginal tooth often reflexed over sorus
Chromosomes: 2n=80
Ecology: Moist or wet rock crevices, talus, cliffs, boulder bases, streamsides
Elevation: 1700–3700 m.
Bioregional distribution: Klamath Ranges, High Cascade Range, High Sierra Nevada, Warner Mountains
Distribution outside California: to Alaska, w Canada, Montana, Colorado; also eastern Canada, Greenland
Synonyms: A. distentifolium Opiz var. a (Butters) Boivin
Var. alpestre in Eurasia
Horticultural information: WET: 4, 5 &SHD: 1, 2, 3, 6, 7, 15, 16, 17; DFCLT.

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bioregional map for ATHYRIUM%20alpestre%20var.%20americanum being generated

Retrieve Jepson Interchange Index to Plant Names entry for Athyrium alpestre var. americanum
Retrieve dichotomous key for Athyrium
Overlay Consortium of California Herbaria specimen data by county on this map
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