Jepson Flora Project: Jepson Interchange    

 
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Jepson Interchange (more information)
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DRYOPTERIDACEAE

WOOD FERN FAMILY

Alan R. Smith and Thomas Lemieux

Perennial, in soil or rock crevices; rhizome generally short-creeping, suberect, or erect, scales large, generally tan to brown, generally 1-colored
Leaves generally tufted, 5–200+ cm, generally ± alike; petiole generally firm, base generally darker, with 2–many vascular strands; blade 1–4-pinnate, often with scales, hair-like scales, hairs (except clear, needle-like hairs generally 0), or short-stalked glands on axes, sometimes between veins, veins free to netted; 1° and 2° axes generally grooved on upper side
Sporangia: sori round, less often oblong or J-shaped, along or at tips of veins; indusia peltate, round-reniform, oblong to linear, J-shaped, hood-like, or cup-like, rarely 0; spores elliptic, winged, ridged, or spiny, scar linear
Genera in family: ± 60 genera, > 1000 species: worldwide, especially tropical, wooded areas. Woodsia sometimes in Woodsiaceae; Athyrium, Cystopteris sometimes in Athyriaceae.

ATHYRIUM

LADY FERN

Rhizome short-creeping to suberect, stout
Leaf: petiole stout, fleshy, easily crushed, straw-colored except base generally blackened, base scaly, in X -section with 2 crescent-shaped vascular strands; blade generally 2-pinnate or more, ± glabrous, veins free
Sporangia: sori ± round, ± oblong, or J-shaped; indusium oblong or J-shaped, laterally attached, or 0
Species in genus: ± 100 species: generally n temp, especially e Asia
Etymology: (Greek: doorless, from enclosed sori)

Native

A. alpestre (D.H. Hoppe) Clairv. var. americanum Butters


Leaf: blade narrow-elliptic or lanceolate, 2–3-pinnate, segments shallowly to deeply pinnately lobed, marginal tooth often reflexed over sorus
Chromosomes: 2n=80
Ecology: Moist or wet rock crevices, talus, cliffs, boulder bases, streamsides
Elevation: 1700–3700 m.
Bioregional distribution: Klamath Ranges, High Cascade Range, High Sierra Nevada, Warner Mountains
Distribution outside California: to Alaska, w Canada, Montana, Colorado; also eastern Canada, Greenland
Synonyms: A. distentifolium Opiz var. a (Butters) Boivin
Var. alpestre in Eurasia
Horticultural information: WET: 4, 5 &SHD: 1, 2, 3, 6, 7, 15, 16, 17; DFCLT.

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