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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

CUPRESSACEAE

CYPRESS FAMILY

Jim A. Bartel

Shrub, tree, evergreen, monoecious or dioecious
Leaves cauline, opposite and 4-ranked or whorled in 3's and 6-ranked, generally scale-like, decurrent, completely covering young stems
Pollen cone small, axillary or terminal
Seed cone ± fleshy to woody, generally hard at maturity; scales opposite or whorled
Seeds 1–many per scale, generally angled or winged, generally wind-dispersed
Genera in family: 17 genera, ± 120 species: worldwide; all North America genera cultivated. [Elias 1980 Complete Trees North America] Juvenile leaves needle- or awl-like, sometimes present in ± mature plants, especially in response to grazing or infection, especially in Cupressus, Juniperus.

CUPRESSUS

CYPRESS

Large shrub, tree, often pyramidal in youth, monoecious
Stem: young shoots generally cylindric (sometimes 4-angled or flat), generally arrayed in 3-dimensional clusters
Leaves opposite, 4-ranked, scale-like, closely appressed, overlapping
Pollen cone generally yellow
Seed cone 6–50 mm, woody, ± spheric to widely cylindric, maturing 1st or 2nd year, often closed > 2 years; scales 6–12, peltate, abutting, shield- or wedge-shaped; projection often present, small, pointed, generally less visible in age
Seeds 2–many per scale, flat, winged; cotyledons 2–5
Chromosomes: 2n=22 for all reports
Species in genus: ± 22 species: w North America, Eurasia
Etymology: (Latin: cypress)
Reference: [Wolf 1948 Aliso 1:1–250]

Native

C. arizonica Greene

Tree 5–25 m
Stem: bark partially peeling in thin strips or plates to fibrous (especially in age), smooth to furrowed, cherry-red to brown to gray; youngest shoots 1–2 mm diam, 4-sided
Leaf dusty gray-green, glaucous or not, appearing blue-green, glandular, often resin-covered
Pollen cone 2–5 mm, 2 mm diam, ± cylindric to 4-sided; scales 8–16; pollen sacs 3–6 per scale
Seed cone 10–35 mm, spheric to ovoid, often warty, dull gray to brown; scales 4–8, projection 0–4 mm
Seed 3–8 mm, light tan to dark brown, slightly warty, often glaucous; attachment scar conspicuous or not
Ecology: Pinyon/juniper or oak/pine woodland, chaparral
Elevation: 750–1800 m.
Bioregional distribution: s Sierra Nevada (Kern, Tulare cos.), Peninsular Ranges
Distribution outside California: Arizona, n Baja California, n&c Mexico

Native

subsp. nevadensis (Abrams) E. Murray

PIUTE CYPRESS

Tree 5–25 m
Stem: bark fibrous, not peeling, gray- to red-brown to cherry-red
Leaf: glands active, conspicuous
Pollen cone: pollen sacs 5–6 per scale
Seed cone 15–35 mm, ovoid, often silver-gray in age; scale projection ± 0
Seed 3–6 mm, light brown, generally glaucous
Chromosomes: 2n=22(23)
Ecology: Pinyon/juniper or oak/pine woodland, chaparral
Elevation: 750–1800 m.
Bioregional distribution: s Sierra Nevada (Kern, Tulare cos.)
Synonyms: C. n. Abrams