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Vascular Plants of California
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Scleranthus annuus subsp. annuus
KNAWEL


Higher Taxonomy
Family: CaryophyllaceaeView DescriptionDichotomous Key
Common Name: PINK FAMILY
Habit: Annual to perennial herb; rarely dioecious (Silene), taprooted or rhizome generally slender. Leaf: simple, generally opposite (subwhorled), entire, pairs at nodes often +- connected at bases; stipules generally 0; petiole generally 0. Inflorescence: generally cyme, generally open; flowers 1--many; involucre generally 0 (present in most Dianthus, Petrorhagia). Flower: generally bisexual, radial; hypanthium often present but obscure; sepals (4)5, +- free or fused into a tube, margins generally scarious, more so on inner 2 or not, tube generally not scarious, awns generally 0; petals (4)5 or 0, generally tapered to base (or with claw long, limb expanded), entire to 2--several-lobed, limb generally without scale-like appendages adaxially, generally without ear-like lobes at base; stamens generally 10, generally fertile, generally free, generally from ovary base; nectaries 0 or 5; ovary superior, generally 1-chambered, placentas basal or free-central, styles 2--5 with 0 branches or 1 with 2--3 branches. Fruit: capsule or utricle (rarely +- dehiscent), generally sessile. Seed: appendage generally 0 (present in Moehringia).
Genera In Family: +- 100 genera, 3000 species: widespread, especially arctic, alpine, temperate northern hemisphere; some cultivated (Agrostemma, Arenaria, Atocion, Cerastium, Dianthus, Gypsophila, Lychnis, Sagina, Saponaria, Silene). Note: Apetalous Caryophyllaceae can also be keyed in Rabeler & Hartman 2005 FNANM 5:5--8. Taxa of Minuartia in TJM2 treated here in Cherleria and Sabulina; Pseudostellaria in Hartmaniella and Torreyostellaria; Vaccaria in Gypsophila; Velezia in Dianthus.
eFlora Treatment Author: Ronald L. Hartman (deceased) & Richard K. Rabeler, except as noted
Scientific Editor: Bruce G. Baldwin & Thomas J. Rosatti.
Genus: ScleranthusView Description 


Habit: Annual, biennial, prostrate to erect, taprooted. Leaf: blade needle-like; vein 1. Inflorescence: axillary, dense; flowers 1--5, +- sessile. Flower: hypanthium widely obovate to urn-shaped, abruptly expanded distally, in fruit hard; sepals 5, free, 1.5--4 mm, narrowly triangular to awl-shaped, glabrous, margin thinly scarious; petals 0; stamens 2--10, on hypanthium rim; styles 2. Fruit: utricle, ovoid. Seed: 1.
Etymology: (Greek: hard flower, from hypanthium in fruit)
eFlora Treatment Author: Ronald L. Hartman (deceased) & Richard K. Rabeler
Reference: Thieret & Rabeler 2005 FNANM 5:149--151
Scleranthus annuus L. subsp. annuus
NATURALIZED
Stem: prostrate to erect, much-branched, generally 4--20 cm, rigid; hairs +- in lines, fine, recurved. Leaf: 4--20+ mm; sheath scarious, ciliate; tip sharp-pointed. Inflorescence: 3--15 mm diam. Flower: 3--4.2 mm; hypanthium 10-ribbed; sepals narrowly triangular to awl-shaped, erect to spreading; styles +- 0.8--1 mm. Seed: 1.4--1.6 mm, widely ovoid, tan except red crescent near acute tip. Chromosomes: 2n=44.
Ecology: Meadows, stream margins, serpentine areas, disturbed areas; Elevation: 300--1200 m. Bioregional Distribution: KR, NCoRO, NCoRH, CaR, n SN, c SNF, PR; Distribution Outside California: to British Columbia, Saskatchewan, eastern North America; native to Europe. Flowering Time: All year
Unabridged Note: 7 other subspecies, native to Europe, western Asia, northern Africa.
Jepson eFlora Author: Ronald L. Hartman (deceased) & Richard K. Rabeler
Reference: Thieret & Rabeler 2005 FNANM 5:149--151
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Botanical illustration including Scleranthus annuus subsp. annuus

botanical illustration including Scleranthus annuus subsp. annuus

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Citation for this treatment: Ronald L. Hartman (deceased) & Richard K. Rabeler 2012, Scleranthus annuus subsp. annuus, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=77174, accessed on April 18, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 18, 2024.

Scleranthus annuus subsp. annuus
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©2005 Steve Matson
Scleranthus annuus subsp. annuus
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©2008 Keir Morse
Scleranthus annuus subsp. annuus
click for enlargement
©2008 Steve Matson
Scleranthus annuus subsp. annuus
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©2020 Neal Kramer
Scleranthus annuus subsp. annuus
click for enlargement
©2005 Steve Matson

More photos of Scleranthus annuus subsp. annuus
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Geographic subdivisions for Scleranthus annuus subsp. annuus:
KR, NCoRO, NCoRH, CaR, n SN, c SNF, PR
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).