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Vascular Plants of California
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Polystichum californicum


Higher Taxonomy
Family: DryopteridaceaeView DescriptionDichotomous Key
Common Name: WOOD FERN FAMILY
Habit: Perennial herb, in soil or rock crevices; rhizome generally short-creeping, suberect, or erect, scales large, generally tan to brown, generally uniformly colored. Leaf: generally tufted, 5--200+ cm, generally +- alike; stipe generally firm, base generally darker, with many vascular strands; blade 1--4-pinnate, often with scales, hair-like scales, hairs (except clear, needle-like hairs generally 0), or short-stalked glands on axes, between veins or not, veins free to netted; rachis, costa generally grooved adaxially. Sporangia: sori round, along veins; indusia peltate or round-reniform; spores elliptic, winged, ridged, or spiny, scar linear.
Genera In Family: +- 40--45 genera, > 1600 species: worldwide, especially tropics, wooded areas. Note: Based on molecular sequence data, Athyrium, Cystopteris, Woodsia removed to Woodsiaceae to preserve a monophyletic Dryopteridaceae.
Unabridged Note: Current data (as reported by Schuettpelz & Pryer and in papers cited therein) suggest that Wooodsiaceae is paraphyletic (with respect to Aspeniaceae, Blechnaceae, and Thelypteridaceae), yet are insufficient to resolve the questions of circumscription (too few taxa, not enough genes sampled). Alternative classifications that would preserve monophyly include recognition of several additional, small families (e.g., Cystopteridaceae, Athyriaceae, and others not in California, each comprising just a few genera) or lumping at least 4 currently recognized families, many of long-standing use and acceptance; a conservative and expedient course is taken for now (Smith et al. 2006), pending further work.
eFlora Treatment Author: Alan R. Smith
Scientific Editor: Thomas J. Rosatti.
Genus: PolystichumView DescriptionDichotomous Key


Common Name: SWORD FERN
Habit: Rhizome generally suberect to erect, often stout. Leaf: stipe stout, firm, generally densely scaly, ×-section with many round vascular strands in an arc; blade 1--3-[> 3--]pinnate, proximal pinnae reduced or not, thin to leathery, scaly, veins generally free, rarely +- jointed; pinna bases often wider acroscopically; teeth, generally including bristle-like tips, < 4 mm [or teeth 0]. Sporangia: sori round; indusium peltate [0 or reniform], sinus 0.
Etymology: (Greek: many rows, from rows of sori on type sp.)
Polystichum californicum (D.C. Eaton) Diels
NATIVE
Leaf: 40--100 cm; stipe < 1/2 blade, base scales 3--4 mm wide, lance-ovate; blade lanceolate, 1- to partly 2-pinnate; pinnae generally 4--7 cm. Sporangia: indusium ciliate. Chromosomes: 2n=82,164.
Ecology: Woodland, streambanks, to rocky open slopes; Elevation: < 1100 m. Bioregional Distribution: NCoRI, NCoRO, s CaRF, SN (3 sites), CW (exc SCoRI), SnBr (1 site); Distribution Outside California: to British Columbia. Note: Probably sterile hybrid between Polystichum munitum or Polystichum imbricans and Polystichum dudleyi. Chromosome doubling (2n=164) restores fertility, allows backcrossing to parents.
Jepson eFlora Author: Alan R. Smith
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

Previous taxon: Polystichum
Next taxon: Polystichum dudleyi

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Botanical illustration including Polystichum californicum

botanical illustration including Polystichum californicum

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Citation for this treatment: Alan R. Smith 2012, Polystichum californicum, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=39407, accessed on April 16, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 16, 2024.

Polystichum californicum
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©2014 Dana York
Polystichum californicum
click for enlargement
©2014 Dana York
Polystichum californicum
click for enlargement
©2014 Dana York
Polystichum californicum
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©2012 John Game
Polystichum californicum
click for enlargement
©2014 Dana York

More photos of Polystichum californicum
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Geographic subdivisions for Polystichum californicum:
NCoRI, NCoRO, s CaRF, SN (3 sites), CW (exc SCoRI), SnBr (1 site)
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).