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Castilleja ambigua subsp. humboldtiensis
HUMBOLDT BAY OWL'S-CLOVER


Higher Taxonomy
Family: OrobanchaceaeView DescriptionDichotomous Key
Common Name: BROOMRAPE FAMILY
Habit: Annual, perennial herb, shrub; often glandular; root-parasites, roots modified into absorptive structures. Stem: generally round in ×-section. Leaf: generally simple, generally alternate, reduced to +- fleshy scales in non-green plants lacking chlorophyll; stipules generally 0. Inflorescence: spike to panicle, generally bracted, or flowers 1--2 in axils. Flower: bisexual; calyx lobes 0--5; corolla generally strongly bilateral, generally 2-lipped (upper lip generally 2-lobed, lower lip generally 3-lobed), abaxial lobes outside other lobes in bud; stamens epipetalous, 4 in 2 pairs (sometimes 1 pair sterile), additional staminode 0(1), anther sacs unequal; ovary superior, chambers 1--2, placentas 2--4, parietal, style 1, stigma lobes 0 or 2. Fruit: capsule, generally +- ovoid, loculicidal, valves 2--4. Seed: many, small, angled; surface smooth or netted.
Genera In Family: +- 100 genera, 2060 species: worldwide, especially northern temperate and Africa. Note: High yield losses in many crops caused by Orobanche and Phelipanche species in Africa, Mediterranean, Middle East, and eastern Europe. Taxa of Orobanche in TJM2 treated here in Aphyllon and Phelipanche.
eFlora Treatment Author: Margriet Wetherwax, except as noted
Scientific Editor: Robert W. Patterson, Bruce G. Baldwin.
Genus: CastillejaView DescriptionDichotomous Key

Common Name: PAINTBRUSH, OWL'S-CLOVER
Habit: Annual to subshrub, green. Leaf: sessile, entire to dissected. Inflorescence: spike-like; bracts becoming shorter, wider, more lobed than leaves, mature tips generally cream to red or green. Flower: calyx unequally 4-lobed, colored like bract tips; corolla upper 2 lip lobes fused, beak-like, tip open, lower lip reduced, 3-toothed to -pouched; stamens 4, anther sacs 2, unequal; stigma entire to 2-lobed, generally exserted. Fruit: +- asymmetric. Seed: generally +- brown, attached at base; coat netted, net-like walls ladder-like or not.
Etymology: (Domingo Castillejo, Spanish botanist, 1744--1793) Note: Hybridization and polyploidy common. Biologically consistent taxa difficult to define. Castilleja chrymactis Pennell not in California, sole (1947) record a misidentified, incomplete specimen.
eFlora Treatment Author: Margriet Wetherwax, T.I. Chuang & Lawrence R. Heckard
Reference: Tank & Olmstead 2008 Amer J Bot 95:608--625
Unabridged Reference: Chuang & Heckard 1991 Syst Bot 16:644--666
Species: Castilleja ambiguaView Description 


Common Name: JOHNNY-NIP
Habit: Annual 10--30 cm, generally much-branched, decumbent, puberulent. Leaf: 10--50 mm, > 3 mm wide, lanceolate to oblong; lobes 0--5. Inflorescence: 3--12 cm, 3--4 cm wide, often dense; bracts 15--25 mm, oblong to ovate, tipped yellow (cream) to rose-purple, lobes 3--9, central lobe generally rounded. Flower: calyx 12--20 mm, divided 1/2 abaxially and on sides, 2/3 adaxially, lobes linear; corolla 14--25 mm, pale yellow or rose-purple, beak 4--5 mm, straight, puberulent, lower lip 3--4 mm, pouches 3--7 mm wide, 1--2 mm deep, generally purple-dotted at base; filaments glabrous; stigma generally +- exserted. Fruit: 8--12 mm. Seed: coat shallowly netted, tight-fitting. Chromosomes: 2n=24.
Note: Highly variable and difficult; many local, +- ecological forms; needs study.
Castilleja ambigua Hook. & Arn. subsp. humboldtiensis (D.D. Keck) T.I. Chuang & Heckard
NATIVE
Habit: Plant +- fleshy, ascending; branches 0--few from mid stem. Inflorescence: bract tips rounded to truncate, +- yellow-pink to rose-purple. Flower: corolla pink or rose-purple (yellow), lower lip teeth 2--3 mm. Seed: 2--2.5 mm.
Ecology: Salt marshes; Elevation: +- 0 m. Bioregional Distribution: NCo (Humboldt, Mendocino cos.), n CCo (Marin Co.). Flowering Time: May--Jun Note: Threatened by coastal development. Yellow-flowered plants from Marin Co. are possible hybrids with Castilleja ambigua subsp. ambigua.
Synonyms: Orthocarpus castillejoides Benth. var. humboldtiensis D.D. Keck
Jepson eFlora Author: Margriet Wetherwax, T.I. Chuang & Lawrence R. Heckard
Reference: Tank & Olmstead 2008 Amer J Bot 95:608--625
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)
Listed on CNPS Rare Plant Inventory

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Botanical illustration including Castilleja ambigua subsp. humboldtiensis

botanical illustration including Castilleja ambigua subsp. humboldtiensis

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Citation for this treatment: Margriet Wetherwax, T.I. Chuang & Lawrence R. Heckard 2012, Castilleja ambigua subsp. humboldtiensis, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=49733, accessed on April 16, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 16, 2024.

Castilleja ambigua subsp. humboldtiensis
click for enlargement
©2017 Steve Matson
Castilleja ambigua subsp. humboldtiensis
click for enlargement
©2017 Steve Matson
Castilleja ambigua subsp. humboldtiensis
click for enlargement
©2017 Steve Matson
Castilleja ambigua subsp. humboldtiensis
click for enlargement
©2017 Steve Matson
Castilleja ambigua subsp. humboldtiensis
click for enlargement
©2017 Steve Matson

More photos of Castilleja ambigua subsp. humboldtiensis
in CalPhotos



Geographic subdivisions for Castilleja ambigua subsp. humboldtiensis:
NCo (Humboldt, Mendocino cos.), n CCo (Marin Co.).
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).