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ONAGRACEAE EVENING-PRIMROSE FAMILY

Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner, except as noted

Annual to perennial herb (to tree). Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous. Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted. Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2–7); petals 4(2–7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric. Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like). Seed: sometimes winged or hair-tufted.
22 genera, ± 657 species: worldwide, especially western North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in northern California. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Unabridged references: [Munz 1965 North America Fl II 5:1–278]

Key to Onagraceae

GAYOPHYTUM

Harlan Lewis

Annual. Stem: generally erect, < 1 m, slender; hairs 0 to dense, rarely glandular. Leaf: cauline, alternate (or ± opposite near base), petioled or not, narrow-lanceolate, entire. Inflorescence: flowers axillary, pedicelled or not, opening at dawn. Flower: hypanthium inconspicuous; sepals 4, staying fused in 2s or all coming free; petals 4, 0.5–8 mm, white, with 1–2 yellow or ± green spots at base, fading pink or red; stamens 8, those opposite sepals longer, pollen ± yellow; ovary chambers 2, stigma generally not exserted beyond anthers, generally touching them, generally ± spheric. Fruit: capsule, ± cylindric or flat; valves 4, generally all coming free, generally equal. Seed: few to many, generally all maturing, generally appressed to septum, alternate or ± opposite between chambers, in each chamber generally in 1 row and generally not overlapped, 0.5–2.3 mm, ovoid, glabrous or hairy, brown or gray mottled with brown; appendages 0.
± 9 species: western North America, 2 South America. (C. Gay, French author of Flora of Chile, 1800–1873) Self-fertile; taxa with petals < 3 mm self-pollinated.
Unabridged references: [Lewis & Szweykowski 1964 Brittonia 16:343–391]

Key to Gayophytum

Previous taxon: Eulobus californicus
Next taxon: Gayophytum decipiens

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 25 2014
Citation for this treatment: [Author of taxon treatment] 2013. Gayophytum, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=9764, accessed on Oct 25 2014

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