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BORAGINACEAE BORAGE or WATERLEAF FAMILY

Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted

Annual to shrub, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1–2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages 0 or 5 at top of tube, alternate stamens; stamens epipetalous; ovary superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1–4, free (fused), smooth to roughened, prickly or bristly or not.
± 120 genera, ± 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum). Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. [Olmstead et al. 2000 Molec Phylogen Evol 16:96–112] Recently treated to include Hydrophyllaceae, Lennoaceae. —Scientific Editors: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.

Key to Boraginaceae

NAMA PURPLE MAT

Sarah Taylor

Perennial herb or generally annual, hairy. Leaf: cauline, generally alternate, margin crenate-dentate or generally entire, wavy, flat, or rolled under. Inflorescence: heads or generally clusters, terminal, not coiled, or flowers 1–2 in leaf axils; bracts 0 or generally leaf-like. Flower: calyx generally free from ovary; corolla salverform to bell-shaped, occasionally cylindric; stamens generally attached to corolla at different levels, generally unequal, part fused to corolla generally narrow-winged, scales at base 0; styles 1–2. Fruit: generally loculicidal, ovoid to elliptic. Seed: generally many, small, red-brown, brown, black, or yellow to orange.
± 55 species: southwestern United States, tropical America, Hawaii. (Greek: a stream) Nama lobbii A. Gray now in Eriodictyon.
Unabridged references: [Ferguson 1998 Syst Bot 23:253–268; Hitchcock 1933 Amer J Bot 20:415–430, 518–534]

Key to Nama

Previous taxon: Myosotis verna
Next taxon: Nama aretioides

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Nov 22 2014
Citation for this treatment: [Author of taxon treatment] 2013. Nama, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=9653, accessed on Nov 22 2014

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