Common Name: WATERLEAF FAMILY Habit: Annual, perennial herb, generally hairy, generally taprooted. Stem: prostrate to erect. Leaf: simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0. Inflorescence: cyme, generally raceme-like and coiled, or flowers 1. Flower: bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla rotate to cylindric, generally deciduous, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chambers 1 or 2, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2--5-chambered, style 1, style branches 2, stigmas generally head-like. Fruit: capsule, generally loculicidal; valves generally 2. Genera In Family: 12 genera, 240--260 species: especially western US; some cultivated (Emmenanthe, Nemophila, Phacelia). Note: Included in Boraginaceae in TJM2 and some other treatments (Angiosperm Phylogeny Group IV 2016 Bot J Linn Soc 181:1--20) but treated as separate family (excluding Namaceae) by Boraginales Working Group (Luebert et al. 2016). eFlora Treatment Author: Genevieve K. Walden, Robert W. Patterson & Richard R. Halse, except as specified Scientific Editor: Bruce G. Baldwin
Etymology: Howell flower, for J.T. Howell, California botanist, 1903--1994 eFlora Treatment Author: Genevieve K. Walden, Robert W. Patterson, Laura M. Garrison & Debra R. Hansen Reference: Walden & Patterson 2010 Madroño 57:268--273.
Howellanthus dalesianus (J.T. Howell) Walden & R. Patt.
NATIVE Habit: Perennial herb 5--15(20) cm, from +- thick caudex. Leaf: in basal rosette, blade 10--50 mm, +- = petiole, oblong to elliptic, entire. Inflorescence: peduncles axillary, from base, few, decumbent, densely glandular-hairy; leaf-like bracts 0--3, subtending inflorescence; flowers 1--several; pedicel 10--20 mm in fruit. Flower: calyx lobes 3--5 mm, 0.5--1 mm wide in flower, 4--7 mm, 1--1.5 mm wide in fruit, +- not alike, especially in fruit, lanceolate to oblanceolate; corolla 5--10 mm, rotate, white, throat purple-marked, limb 5--15 mm diam, corolla scales free from filaments, semi-circular; stamens (4)6--8 mm, included to slightly exserted, equal, attached at same level, glabrous, filament not widened at base; style (3)6--7 mm, +- included, cleft nearly to base, generally glabrous, occasionally sparsely hairy proximally. Fruit: +- 4 mm, +- spheric, hairy. Seed: 2--4, 2.5--4 mm; surface honeycombed or pitted, attached fleshy structure 0. Chromosomes: n=8. Ecology: Ultramafic substrates, meadows, streambanks, conifer forest; Elevation: 1500--2000 m. Bioregional Distribution: KR. Flowering Time: May--Aug Synonyms: Phacelia dalesiana J.T. Howell Jepson eFlora Author: Genevieve K. Walden, Robert W. Patterson, Laura M. Garrison & Debra R. Hansen Reference: Walden & Patterson 2010 Madroño 57:268--273. Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Previous taxon: Howellanthus Next taxon: Hydrophyllum
Botanical illustration including Howellanthus dalesianus
Citation for this treatment: Genevieve K. Walden, Robert W. Patterson, Laura M. Garrison & Debra R. Hansen 2021, Howellanthus dalesianus, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=95351, accessed on April 18, 2024.
Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 18, 2024.
MAP CONTROLS 1. You can change the display of the base map layer control box in the upper right-hand corner.
2. County and Jepson Region polygons can be turned off and on using the check boxes.
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
Data provided by the participants of the
Consortium of California Herbaria.
MAP LEGEND View all CCH records All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
READ ABOUT YELLOW FLAGS
CCH collections by month
Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).
