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CERATOPHYLLACEAE HORNWORT FAMILY

Donald H. Les

Perennial herb, submersed aquatic; overwintering on bottom as detached, dense shoot tips (winter buds); roots 0; monoecious; water-pollinated. Stem: flexible with water currents, internodes clustered near tip. Leaf: whorled, generally compound, forked; segments linear, toothed; stipules 0; petiole 1–2 mm, translucent. Inflorescence: reduced axillary cymes generally near shoot tips, ± sessile, 1–several per node, generally staminate below, pistillate above, or sometimes mixed at a node; flowers 1–3 per cyme but appearing to be 1 subtended by calyx-like whorl of 8–15 bracts, bracts linear, leaf-like, fused at base, toothed at tip, persistent in fruit. Flower: unisexual; perianth 0; staminate round-topped, stamens 3–50, spirally arranged, anthers ± sessile, exserted from bracts, pollen shed underwater; pistillate of 1 simple, ovoid pistil, ovary superior, chamber 1; placenta pendulous; style elongate, spine-like, with decurrent groove; stigma pocket-like, at style base; ovule 1. Fruit: achene, tubercled or not, spines 0 or elongate, basal, facial, marginal or stylar; lateral margin winged or not.
1 genus, 6 species: ± worldwide. [Les 1997 FNANM 3:81–84] —Scientific Editor: Thomas J. Rosatti.

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Jul 31 2014
Citation for this treatment: [Author of taxon treatment] 2013. Ceratophyllaceae, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=94, accessed on Jul 31 2014

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