Jepson Herbarium
		University of California, Berkeley

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Tree, generally monoecious, wind-pollinated; hairs many-branched.
Stem: bark peeling in scaly plates, leaving ± smooth areas of various colors, in age dark, thick, fissured; twig hairs dense.
Leaf: simple, alternate, deciduous; lobes, veins generally 3,5(7), palmate; stipules generally leaf-like, free or fused around stem, shed by leaf maturity or not; petiole at base dilated, hollow, ± covering bud; blade hairs dense, ± 0 in age.
Inflorescence: heads 1–7, ± evenly spaced on axis, spheric, many-flowered, sessile or on pendent peduncles, generally unisexual; staminate breaking apart in age; pistillate persistent; bracts subtending heads, flowers.
Flower: unisexual; calyx cup-shaped, sepals (0)3–6(8), free or united basally.
Staminate flower: petals 3–6, minute or vestigial, fleshy or scale-like; stamens 3–6(8), alternate petals, anthers subsessile, axis above anther expanded, disk-like, ± peltate; carpels vestigial or 0.
Pistillate flower: petals 3–6, minute, or generally 0; staminodes often 3–4; carpels (3)5–9, free, ovary of each superior, 1-chambered, generally 1-ovuled, with 1 ± linear style.
Fruit: achenes in spheric head, small, each with hairs from base, shorter hairs up the side; style persistent, beak-like, or deciduous.
1 genus, ± 8 species: n temperate; some cultivated for ornamental, shade; wood generally of limited commercial value. [Feng et al. 2005 Syst Bot 30:786–799] —Scientific Editor: Thomas J. Rosatti.
Unabridged genera in family: 1 genus, ± 8 species: n temp; some cult for ornamental, shade; wood generally of limited commercial value, although long ago used for buttons, the trees then called buttonwood.
Unabridged note: Lvs and twigs with many-branched hairs, each comprising a single multicellular central axis with unicellular lateral, generally whorled rays.

(Greek: probably broad, for leaves) Fr length excludes style.

Stem: 12–24(36) m, erect; base generally <<= 0.5 m wide; outer bark light gray, inner green-cream.
Leaf: stipules (0)1–2 cm, generally deciduous by maturity; petiole 2–10 cm; blade ± 11–18 cm, ± round or elliptic, ± glabrous.
Staminate flower: calyx reduced, cup-like; petals free.
Pistillate flower: calyx reduced, cup-like.
Fruit: head 2.5–3.5 cm, generally sessile; achene 7.5–11 mm, top truncate, basal hairs 1/2–2/3 fruit length, persistent on fruit, head; style deciduous.
In cultivation, may persist from old homesteads, escapes to waterways; 16–155 m. Cascade Range Foothills, n Sierra Nevada Foothills, Sacramento Valley;

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occurs	Red area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon; markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
Chart based on elevation range in Manual and elevations and coordinates of CCH records. Data provided by the participants of the Consortium of California Herbaria. Note: About half of the CCH records include both elevation and coordinates.	Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria. View all CCH records   CCH collections by month Duplicates counted once; synonyms included. Species do not include records of infraspecific taxa. Blue line denotes Manual flowering time.