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Perennial herb from caudex or rhizome, generally ± hairy. Stem: often ± leafy on proximal 1/2, rarely trailing and leafy throughout. Leaf: generally simple, basal and/or cauline, generally alternate, generally petioled; veins ± palmate. Inflorescence: raceme or panicle, generally ± scapose. Flower: generally bisexual, generally radial; hypanthium free to ± fused to ovary; calyx lobes generally 5; petals generally 5, free, generally clawed, generally white; stamens 3, , 5, 8, or 10; pistils 1 (carpels ± fused, ovary lobed, chambers 1 or 2, placentas generally 2(3), axile or parietal or occasionally proximally axile and distally marginal in ovary lobes) or 2 (carpels free, placentas marginal), ovary nearly superior to inferior, occasionally more superior in fruit, styles generally 2(3). Fruit: capsule (generally 2(3)-beaked, valves generally 2(3), generally equal) or 2 follicles. Seed: generally many, small.
± 30 genera, 600 species: especially northern temperate, arctic, alpine; some cultivated (Bergenia, Darmera, Heuchera, Saxifraga, Tellima, Tolmiea). [Soltis et al. 2001 Ann Missouri Bot Gard 88:669–693; Okuyama et al. 2008 Molec Phylogen Evol 46:560–575] California Mitella moved to Mitellastra, Ozomelis, Pectiantia; Suksdorfia ranunculifolia to Hemieva. Parnassia moved to Parnassiaceae. —Scientific Editor: Bruce G. Baldwin.
Unabridged references: [Soltis 1988 Syst Bot 13:64–72]
Key to Saxifragaceae
Plant generally ± hairy, often glandular; caudex or rhizome generally not woody, generally scaly. Leaf: basal (cauline); blade linear to (ob)ovate or ± round, base tapered to reniform, margin entire or toothed. Inflorescence: flowers few to many; bracts scale-like. Flower: generally radial; hypanthium free or ± fused to ovary; petals 5, white, sometimes with yellow spots at base; stamens 10, filaments flat or variously inflated; pistils 1 (chambers 2, placentas 2, axile or occasionally proximally axile and distally marginal) or 2, ovary superior to ± inferior (sometimes more superior in fruit), styles free throughout. Fruit: capsule or 2 follicles.Key to Micranthes
± 80 species: North America, Eurasia, South America, especially cool temperate northern hemisphere. (Latin: small flower) [Elvander 1984 Syst Bot Monogr 3:1–44] Intermediates common between Micranthes integrifolia, Micranthes nidifica, Micranthes fragosa, Micranthes aprica; some may be vegetatively reproducing, sterile hybrids. Study needed.
Unabridged references: [Small & Rydberg 1905 N Am Fl 22:132–150]
Plant 15–35 cm; caudex generally with rhizomes and bulblets. Leaf: 4–11 cm; petiole 2–5 cm; blade triangular to ovate, base tapered, entire to minutely toothed. Inflorescence: open to ± congested at tips, branches spreading to ascending; pedicel densely glandular. Flower: sepals ± reflexed to spreading, < petals; petals 2–3 mm, obovate; filaments ± flat, narrowed near tip; nectaries disk-like, lobed; pistils 2, ovary >> 1/2-inferior in flower. Fruit: 2 follicles.
2n=20,38. Moist, rocky slopes and outcrops; 500–2000 m. Klamath Ranges, Cascade Range, Modoc Plateau (except Warner Mountains). [Saxifraga nidifica Greene var. claytoniifolia (Small) Elvander] Intermediate between Micranthes californica and Micranthes integrifolia. Plants with 2n=38 chromosomes possibly best treated as variant of Micranthes integrifolia. Mar–Jun [Online Interchange]
Unabridged synonyms: [Saxifraga nidifica Greene var. claytoniifolia (Canby ex Small) Elvander]
Unabridged note: Expanded author citation: Micranthes fragosa (Suksd. ex Small) Small
Previous taxon: Micranthes ferruginea
Next taxon: Micranthes howellii
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Dec 20 2014
Citation for this treatment: [Author of taxon treatment] 2013. Micranthes, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=91945, accessed on Dec 20 2014
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|Geographic subdivisions indicated for the distribution of Micranthes fragosa|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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