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SAXIFRAGACEAE SAXIFRAGE FAMILY

Michael S. Park & Patrick E. Elvander, except as noted

Perennial herb from caudex or rhizome, generally ± hairy. Stem: often ± leafy on proximal 1/2, rarely trailing and leafy throughout. Leaf: generally simple, basal and/or cauline, generally alternate, generally petioled; veins ± palmate. Inflorescence: raceme or panicle, generally ± scapose. Flower: generally bisexual, generally radial; hypanthium free to ± fused to ovary; calyx lobes generally 5; petals generally 5, free, generally clawed, generally white; stamens 3, [4], 5, 8, or 10; pistils 1 (carpels ± fused, ovary lobed, chambers 1 or 2, placentas generally 2(3), axile or parietal or occasionally proximally axile and distally marginal in ovary lobes) or 2 (carpels free, placentas marginal), ovary nearly superior to inferior, occasionally more superior in fruit, styles generally 2(3). Fruit: capsule (generally 2(3)-beaked, valves generally 2(3), generally equal) or 2 follicles. Seed: generally many, small.
± 30 genera, 600 species: especially northern temperate, arctic, alpine; some cultivated (Bergenia, Darmera, Heuchera, Saxifraga, Tellima, Tolmiea). [Soltis et al. 2001 Ann Missouri Bot Gard 88:669–693; Okuyama et al. 2008 Molec Phylogen Evol 46:560–575] California Mitella moved to Mitellastra, Ozomelis, Pectiantia; Suksdorfia ranunculifolia to Hemieva. Parnassia moved to Parnassiaceae. —Scientific Editor: Bruce G. Baldwin.
Unabridged references: [Soltis 1988 Syst Bot 13:64–72]

Key to Saxifragaceae

MICRANTHES SAXIFRAGE

Michael S. Park

Plant generally ± hairy, often glandular; caudex or rhizome generally not woody, generally scaly. Leaf: basal (cauline); blade linear to (ob)ovate or ± round, base tapered to reniform, margin entire or toothed. Inflorescence: flowers few to many; bracts scale-like. Flower: generally radial; hypanthium free or ± fused to ovary; petals 5, white, sometimes with yellow spots at base; stamens 10, filaments flat or variously inflated; pistils 1 (chambers 2, placentas 2, axile or occasionally proximally axile and distally marginal) or 2, ovary superior to ± inferior (sometimes more superior in fruit), styles free throughout. Fruit: capsule or 2 follicles.
± 80 species: North America, Eurasia, South America, especially cool temperate northern hemisphere. (Latin: small flower) [Elvander 1984 Syst Bot Monogr 3:1–44] Intermediates common between Micranthes integrifolia, Micranthes nidifica, Micranthes fragosa, Micranthes aprica; some may be vegetatively reproducing, sterile hybrids. Study needed.
Unabridged references: [Small & Rydberg 1905 N Am Fl 22:132–150]

Key to Micranthes

M. californica (Greene) Small
NATIVE
Plant 15–35 cm; caudex generally with long slender rhizomes and bulblets. Leaf: 4–10 cm; petiole 2–5 cm; blade ovate, base tapered, teeth shallow, ± round or sharp. Inflorescence: often 1-sided, ± open. Flower: sepals generally reflexed, elliptic; petals 2.5–4.5 mm, >> sepals, elliptic to round; filaments ± flat, generally narrowed at tip; nectaries disk-like, lobed; pistils 2, ovary ± 1/2-inferior in flower. Fruit: 2 follicles.
2n=20. Moist, shady places; < 1200 m. Klamath Ranges, Outer North Coast Ranges, Inner North Coast Ranges, Cascade Range Foothills, Sierra Nevada (expected Tehachapi Mountain Area), Great Central Valley (Sutter to Stanislaus cos.), Central Western California, Southwestern California; southwestern Oregon, northwestern Baja California. [Saxifraga californica Greene] Plants in far northern California difficult to assign; see Micranthes fragosa. Feb–May(Jun) [Online Interchange]

Previous taxon: Micranthes bryophora
Next taxon: Micranthes ferruginea

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 28 2014
Citation for this treatment: [Author of taxon treatment] 2013. Micranthes, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=91942, accessed on Aug 28 2014

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Geographic subdivisions indicated for the distribution of Micranthes californica Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.