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ONAGRACEAE EVENING-PRIMROSE FAMILY

Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner, except as noted

Annual to perennial herb (to tree). Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous. Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted. Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2–7); petals 4(2–7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric. Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like). Seed: sometimes winged or hair-tufted.
22 genera, ± 657 species: worldwide, especially western North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in northern California. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Unabridged references: [Munz 1965 North America Fl II 5:1–278]

Key to Onagraceae

NEOHOLMGRENIA
Annual from taproot; branches many, slender, ascending from base. Stem: nearly naked proximally, densely leafy distally. Leaf: cauline, alternate, mostly closely spaced at stem tips, entire. Inflorescence: flowers 1 in axils. Flower: opening at dawn; sepals (3)4, reflexed separately or in pairs; petals (3)4, yellow, fading yellow; stamens (4)8, longer ones opposite sepals, anthers attached at middle, pollen grains 3-angled except in polyploid taxa (visible with hand lens); stigma hemispheric, > anthers and cross-pollinated or ± = anthers and self-pollinated. Fruit: sessile, 0.5–1 cm, ± straight, strongly flattened, walls ± swollen by seeds. Seed: in 1 row per chamber, narrowly obovoid, smooth, shiny, dots or blotches 0.
2 species: northeastern California, Nevada, central Utah, British Columbia, Alberta. (Arthur H. Holmgren (1912–1992), Noel H. Holmgren (b. 1937), and Patricia K. Holmgren (b. 1940)) [Wagner & Hoch 2009 Novon 19:130–132] Incl in Camissonia in TJM (1993).
Unabridged references: [Raven 1969 Contr US Natl Herb 37:161–396; Wagner et al. 2007 Syst Bot Monogr 83:1–240]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 20 2014
Citation for this treatment: [Author of taxon treatment] 2013. Neoholmgrenia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=91930, accessed on Oct 20 2014

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