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CLEOMACEAE SPIDERFLOWER FAMILY

Robert E. Preston & Staria S. Vanderpool

Annual, perennial herb, shrub, often ill-smelling. Leaf: generally 1-palmate, generally alternate, generally petioled; stipules generally minute, often bristle-like or hairy; leaflets 0 or 3–7. Inflorescence: raceme, head, or flowers 1, expanded in fruit; bracts generally 3-parted below, simple above, or 0. Flower: generally bisexual, radial to ± bilateral; sepals generally 4, free or fused, generally persistent; petals generally 4, free, ± clawed; stamens generally 6, free, exserted, anthers generally coiling at dehiscence; ovary superior, generally on stalk-like receptacle, chamber generally 1, placentas generally 2, parietal, style 1, persistent, stigma generally minute, ± head-like. Fruit: 2 nutlets or generally capsule, septicidal; valves generally 2, deciduous, leaving septum (frame-like placentas) behind; pedicel generally ± reflexed to spreading.
17 genera, ± 150 species: widespread tropics to arid temperate. [Iltis & Cochrane 2007 Novon 17:447–451] Treated as Capparaceae in TJM (1993). —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Ernst 1963 J Arnold Arbor 44:81–93]

Key to Cleomaceae

PERITOMA BEE PLANT
Annual, shrub, glabrous to densely glandular-hairy. Stem: generally branched from upper nodes. Leaf: petiole 5–45 mm; leaflets 3–5. Inflorescence: raceme, terminal, generally 1–4 cm, generally 5–40 cm in fruit; pedicels 4–20 mm. Flower: often ± unisexual (stamens or pistils vestigial), ± bilateral, parts generally yellow; sepals free or fused, deciduous above base; petal claw 0 to short. Fruit: longer than wide; septum linear to oblong; receptacle stalk-like, reflexed to ascending. Seed: 10–40.
6 species. (Greek: cutting all-around, from calyx) [Iltis & Cochrane 2007 Novon 17:447–451]

Key to Peritoma

P. arborea (Nutt.) H.H. Iltis BLADDERPOD
NATIVE
Shrub, much branched, generally 5–20 dm, minutely hairy. Leaf: petiole 1–3 cm; leaflets generally 3, 15–45 mm, oblong-elliptic. Inflorescence: 1–30 cm; pedicels 8–15 mm, thicker in fruit. Flower: sepals fused in basal 1/2, 4–7 mm, ± entire, green; petals 8–14 mm, 4–5 mm wide, yellow, claw 0; stamens 15–25 mm, yellow, anthers 2–2.5 mm; style 0.9–1.2 mm or pistil aborting in bud. Fruit: capsule, 3–6 cm, 1–2.5 cm wide, inflated, smooth, leathery, light brown, tardily dehiscent; valves 2–3; receptacle stalk-like, 1–2 cm, stout, reflexed or not.
2n=34,40. [Isomeris arborea Nutt.] Varieties ± distinct. [Online Interchange]

P. arborea var. arborea
NATIVE
Fruit: obovoid, tapered to receptacle, tapered to tip or abruptly narrowed to short beak.
Slopes, grassland, coastal scrub; 5–1200 m. South Coast, s Channel Islands, Peninsular Ranges; Baja California. [Isomeris arborea var. arborea] All year [Online Interchange]

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Next taxon: Peritoma arborea var. globosa

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 19 2014
Citation for this treatment: [Author of taxon treatment] 2013. Peritoma, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=91923, accessed on Sep 19 2014

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Geographic subdivisions indicated for the distribution of Peritoma arborea var. arborea Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.