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Martin F. Wojciechowski, except as noted

Annual to tree. Leaf: generally alternate, generally compound, generally stipuled, generally entire, pinnately veined Inflorescence: generally raceme, spike, umbel or head; or flowers 1–few in axils. Flower: generally bisexual, generally bilateral; hypanthium 0 or flat to tubular; sepals generally 5, generally fused; petals generally 5, free, fused, or lower 2 ± united into keel (see 3, Key to Groups, for banner, wings); stamens 10 or many (or [1], 5, 6, 7, 9), free or fused or 10 with 9 filaments at least partly fused, 1 (uppermost) free; pistil 1, ovary superior, generally 1-chambered, ovules 1–many, style, stigma 1. Fruit: legume, including a stalk-like base (above receptacle) or not. Seed: 1–many, often ± reniform, generally hard, smooth.
± 730 genera, 19400 species: worldwide; with grasses, requisite in agriculture, most natural ecosystems. Many cultivated, most importantly Arachis, peanut; Glycine, soybean; Phaseolus, beans; Medicago, alfalfa; Trifolium, clovers; many orns. [Lewis et al. (eds) 2005 Legumes of the World. RBG, Kew] Unless stated otherwise, fruit length including stalk-like base, number of 2° leaflets is per 1° leaflet. Upper suture of fruit adaxial, lower abaxial. Anthyllis vulneraria L. evidently a waif, a contaminant of legume seed from Europe. Laburnum anagyroides Medik., collected on Mount St. Helena in 1987, may be naturalized. Ceratonia siliqua L., carob tree (Group 2), differs from Gleditsia triacanthos L. in having evergreen (vs deciduous) leaves that are 1-pinnate (vs 1-pinnate on spurs on old stems, 2-pinnate on new stems) with 2–5(8) (vs 7–17) 1° leaflets, commonly cultivated, now naturalized in southern California. Aeschynomene rudis Benth. <Noxious weed>, Halimodendron halodendron (Pall.) Voss <Noxious weed> (possibly extirpated), Lens culinaris Medik. are agricultural weeds. Caragana arborescens Lam. only cultivated. Ononis alopecuroides L. <Noxious weed>, Sphaerophysa salsula (Pall.) DC. <Noxious weed> all evidently extirpated. Cercidium moved to Parkinsonia; Chamaecytisus to Cytisus; Psoralidium lanceolatum to Ladeania. —Scientific Editors: Martin F. Wojciechowski, Thomas J. Rosatti.

Key to Fabaceae


Luc Brouillet

Annual, perennial herb, shrub, unarmed. Leaf: generally odd-1-pinnate (or ± palmately compound, rarely some or most simple); stipules often gland-like, bump-like, or conic, often not apparent; leaflets 3–9, generally irregularly arranged, lowest not stipular in position. Inflorescence: umbel or 1–2-flowered, axillary, generally peduncled, often bracted. Flower: corolla generally yellow (white, pink), fading darker; 9 filaments fused, 1 free. Fruit: dehiscent or not, exserted from calyx or not, ovoid to oblong, ± beaked. Seed: 1–several.
± 23 species: southwestern Canada, western United States, Mexico, 1 sp. in Chile. (Greek: tip, probably for hooked-tipped fruit) [Brouillet 2008 J Bot Res Inst Texas 2:387–394] Intermediates may be hybrids.
Unabridged etymology: (Greek acme, point, apex, probably for the hooked-tipped fruit)
Unabridged references: [Isely 1981 Mem New York Bot Gard 25:128–206; Sokoloff 2000 Ann Bot Fenn 37:125–131]
Unabridged note: Pollen has 4(7) apertures.

Key to Acmispon

A. dendroideus (Greene) Brouillet
Perennial herb, shrub, sparsely or densely strigose or ± silky, green or gray. Stem: decumbent to ascending, 5–20 dm. Leaf: irregularly pinnate; stipules gland-like; leaflets 7–15 mm, elliptic to obovate. Inflorescence: 3–10-flowered, peduncle 0 or 2–10 mm. Flower: calyx 4–6 mm, lobes < tube, thinly strigose to ± glabrous; corolla 8–12 mm, yellow, wings ± = keel, stigma glabrous. Fruit: indehiscent, divergent to descending, soon-exserted, oblong-tapering, straight or curved, not flat; beak narrow, abruptly curved, 2–3 mm.
2n=14. [Lotus dendroideus (Greene) Greene] Island-to-island variants recognized variously; the following distinct in morphology, geography. [Online Interchange]

A. dendroideus var. dendroideus ISLAND BROOM
Leaf: leaflets 3–5, finely or sparsely strigose, ± green. Inflorescence: peduncle bract 0. Fruit: 1–1.5 cm.
Bluffs, inland canyons, open sites near ocean; < 350 m. Channel Islands (except San Miguel, San Clemente islands). [Lotus dendroideus (Greene) Greene var. dendroideus; Lotus scoparius var. dendroideus (Greene) Ottley] Jan–Aug [Online Interchange] {CNPS list}

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 26 2015
Citation for this treatment: [Author of taxon treatment] 2013. Acmispon, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 26 2015

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Geographic subdivisions indicated for the distribution of Acmispon dendroideus var. dendroideus Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.