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Vascular Plants of California
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Vancouveria
INSIDE-OUT-FLOWER

Higher Taxonomy
Family: BerberidaceaeView DescriptionDichotomous Key

Common Name: BARBERRY FAMILY
Habit: Perennial herb, shrub, [tree], generally rhizomed, caudexed or not, glabrous, glaucous, or hairy. Stem: spreading to erect, branched or not. Leaf: simple, 1--3-ternate, or pinnately compound, basal and cauline, generally alternate, deciduous or evergreen, petioled, stipuled. Inflorescence: generally raceme, spike, or panicle, scapose, terminal, or axillary. Flower: generally bisexual, radial; sepals 6--18 or 0, generally in whorls of 3; petals generally 6, in 2 whorls of 3, or 0; stamens 6--12(13), free or fused at base, in 2 whorls or not, anthers dehiscent by flap-like valves or longitudinal slits; ovary superior, chamber 1, ovules generally 1--10, style 1 or 0, stigma flat or spheric. Fruit: berry, capsule, achene [follicle].
Genera In Family: 16 genera, +- 670 species: temperate, tropics worldwide; some cultivated (Berberis, Epimedium, Nandina (heavenly bamboo), Vancouveria). Note: Lower sepals sometimes called "bracteoles," inner petals "staminodes".
eFlora Treatment Author: Michael P. Williams
Scientific Editor: Thomas J. Rosatti.
Vancouveria
Habit: Perennial herb, rhizomes extensive, scales brown. Stem: 0. Leaf: basal, long-petioled, 2--3-ternate; leaflet blades ovate to +- cordate, lobes 3, shallow, teeth 0 or shallow. Inflorescence: raceme or panicle, +- scapose, open, long-peduncled; flowers spreading to pendent. Flower: sepals generally 12--15, 8--9 mm, outer 6--9 << inner 6, bract-like, deciduous, inner petal-like, persistent, in age reflexed; petals 6, < inner sepals, reflexed from base, distally glandular; stamens generally 6, held against ovary, style, anther valves flap-like, pointed tipward; ovules 2--10, style 1, < ovary, persistent, beak-like in fruit, stigma cup-like. Fruit: capsule, 2-valved, generally elliptic. Seed: with oily body for ant dispersal.
Species In Genus: 3 species: temperate western North America. Etymology: (Captain George Vancouver, British explorer, 1757--1798) Note: Pedicel appears to arise from inside flower, from tip instead of base, yielding an upside-down or "inside-out" flower.
Jepson eFlora Author: Michael P. Williams
Reference: Zhang 2007 Syst Bot 32:81--92
Unabridged Reference: Stearn 1938 J Linn Soc Bot 51:409--535
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)
Key to Vancouveria

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Citation for this treatment: Michael P. Williams 2012, Vancouveria, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=8936, accessed on April 25, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 25, 2024.