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Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner, except as noted

Annual to perennial herb (to tree). Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous. Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted. Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2–7); petals 4(2–7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric. Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like). Seed: sometimes winged or hair-tufted.
22 genera, ± 657 species: worldwide, especially western North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in northern California. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Unabridged references: [Munz 1965 North America Fl II 5:1–278]

Key to Onagraceae


1 sp.: western United States. (Latin: diminutive of Chylismia) [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Incl in Camissonia in TJM (1993).
Unabridged references: [Raven 1969 Contr US Natl Herb 37:161–396]

C. pterosperma (S. Watson) W.L. Wagner & Hoch
Annual, from taproot; hairs bristly, in inflorescence also glandular. Stem: erect, branched, 2–14 cm, slender, peeling near base. Leaf: cauline, alternate, ± sessile; blade 3–30 mm, narrowly lanceolate to oblanceolate, entire. Inflorescence: raceme, nodding in bud. Flower: opening at dawn; hypanthium 1–2 mm; sepals 4, 1.5–2.5 mm, reflexed singly or occasionally in pairs; petals 4, 1.5–2.5 mm, notched at tip, white, with yellow area at base, fading purple; longer stamens opposite sepals, anthers attached at base, pollen grains 3-angled; stigma hemispheric, ± = anthers, self-pollinated. Fruit: (3)4-chambered, ascending or spreading, 12–28 mm, 0.6–0.8 mm wide, straight or slightly curved, cylindric, slightly swollen by seeds; pedicel 4–8 mm. Seed: in 2 rows per chamber, 1–1.5 mm, with concave and convex sides, concave side with a thick wing, both sides densely covered with glassy club-shaped hairs.
2n=14. Uncommon. Well-drained, generally volcanic slopes, pinyon/juniper woodland, sagebrush scrub; 1400–2400 m. White and Inyo Mountains (Inyo Mtns), Desert Mountains (Panamint Range); to Oregon, western Utah, northwestern Arizona. [Camissonia pterosperma (S. Watson) P.H. Raven; Oenothera pterosperma S. Watson] Self-pollinated. May–Jun [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 29 2015
Citation for this treatment: [Author of taxon treatment] 2013. Chylismiella, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 29 2015

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Geographic subdivisions indicated for the distribution of Chylismiella pterosperma Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.