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ONAGRACEAE EVENING-PRIMROSE FAMILY

Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner, except as noted

Annual to perennial herb (to tree). Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous. Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted. Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2–7); petals 4(2–7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric. Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like). Seed: sometimes winged or hair-tufted.
22 genera, ± 657 species: worldwide, especially western North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in northern California. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Unabridged references: [Munz 1965 North America Fl II 5:1–278]

Key to Onagraceae

CAMISSONIOPSIS
Annual to subshrub, from taproot. Leaf: basal and cauline, alternate, simple, generally narrowly lanceolate or narrowly elliptic to ovate. Inflorescence: spike, nodding in bud, generally flower from basal-most to distal nodes. Flower: opening at dawn; sepals 4, reflexed singly or in fused pairs; petals 4, yellow, fading red, generally with 1+ red basal spots, with no ultraviolet pattern; stamens 8, longer opposite sepals, anthers attached at middle, pollen grains 3-angled except in polyploid taxa at 20×; ovary chambers 4, stigma ± spheric or hemispheric, exceeding anthers and cross-pollinated or ± = anthers and self-pollinated. Fruit: 4-angled at least when dry, generally proximally thick, contorted or curled 1–5 times, or straight, not swollen by seeds, sessile. Seed: in 1 row per chamber, narrowly obovoid, flattened, dull brown-black.
14 species: California, Arizona, Oregon, Baja California. (Greek: like Camissonia) [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Polyploidy and self-pollination have predominated in evolution of genus. Incl in Camissonia in TJM (1993).
Unabridged references: [Raven 1969 Contr US Natl Herb 37:161–396]

Key to Camissoniopsis

C. cheiranthifolia (Spreng.) W.L. Wagner & Hoch BEACH EVENING-PRIMROSE
NATIVE
Perennial herb or subshrub, short-lived, rosetted, densely strigose (glabrous); hairs of inflorescence generally erect, short. Stem: prostrate to ± ascending, < 60(130) cm, peeling. Leaf: 5–50 mm, narrowly ovate to obovate, minutely serrate; cauline petioles 0–10 mm. Flower: hypanthium 2.1–8.5 mm; sepals 4–11.5 mm; petals 6–20 mm, basal spots 0–2. Fruit: 10–25 mm, 2–2.5 mm wide, 4-angled, generally 1–2-coiled. Seed: 1.2–1.3 mm.
2n=14. [Camissonia cheiranthifolia (Spreng.) Raim.] Subspecies intergrade on Channel Islands. [Online Interchange]
Unabridged synonyms: [Oenothera cheiranthifolia Hornem. ex Spreng.; Oenothera cheiranthifolia var. nitida (Greene) Munz; Camissonia cheiranthifolia (Hornem. ex Spreng.) Raim.]
Unabridged note: Expanded author citation: Camissoniopsis cheiranthifolia (Hornem. ex Spreng.) W.L. Wagner & Hoch

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 2 2014
Citation for this treatment: [Author of taxon treatment] 2013. Camissoniopsis, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=89197, accessed on Sep 2 2014

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Geographic subdivisions indicated for the distribution of Camissoniopsis cheiranthifolia Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.