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ONAGRACEAE EVENING-PRIMROSE FAMILY

Warren L. Wagner & Peter C. Hoch, family description, key to genera; treatment of genera by Warren L. Wagner, except as noted

Annual to perennial herb (to tree).
Leaf: cauline or basal, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous.
Inflorescence: spike, raceme, panicle, or flowers 1 in axils; bracted.
Flower: generally bisexual, generally radial, often opening at either dawn or dusk; hypanthium generally prolonged beyond ovary (measured from ovary tip to sepal base); sepals 4(2–7); petals 4(2–7, rarely 0), often fading darker; stamens 2 × or = sepals in number, anthers 2-chambered, opening lengthwise, pollen interconnected by threads; ovary inferior, chambers generally as many as sepals (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, spheric, or hemispheric.
Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like).
Seed: sometimes winged or hair-tufted.
22 genera, ± 657 species: worldwide, especially w North America; many cultivated (Clarkia, Epilobium, Fuchsia, Oenothera). [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Gaura moved to Oenothera. Fuchsia magellanica Lam. naturalized in n CA. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.
Unabridged references: [Munz 1965 North America Fl II 5:1–278]

Key to Onagraceae

TETRAPTERON
Annual, stem 0, from taproot.
Leaf: mostly basal, or cauline on short stems, alternate, broadly sessile or clasping.
Inflorescence: flowers 1 in axils of rosette, nodding in bud.
Flower: opening at dawn; sepals 4, reflexed in pairs; petals 4, yellow, unspotted, strongly ultraviolet-reflective, fading red-orange; longer stamens opposite sepals, anthers attached near base, pollen grains 3-angled; ovary tip with sterile 0.4–18 cm projection that often breaks off in fruit, with clear abscission line at juncture between short hypanthium and fertile part of ovary, stigma hemispheric, ± = anthers and generally self-pollinated (or > anthers and cross-pollinating).
Fruit: irregularly obovoid, sharply 4-angled with pointed wing near center top of each valve, ± sessile, very tardily dehiscent in distal 1/2 only, thick-walled (not distended by seeds), persistent for > 1 yr, sessile.
Seed: in 2 crowded rows per chamber, obovoid to narrowly obovoid, finely papillate or deeply pitted, tan or brown with dark patches.
2 species: CA, disjunct in nw NV, s OR, n Baja CA. (Greek: four winged) [Wagner et al. 2007 Syst Bot Monogr 83:1–240] Incl in Camissonia in TJM (1993).
Unabridged references: [Raven 1969 Contr US Natl Herb 37:161–396]

Key to Tetrapteron

Previous taxon: Taraxia tanacetifolia
Next taxon: Tetrapteron graciliflorum

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
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map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
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