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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central North Coast Ranges, Carrichtera annua (L.) DC. in South Coast, Iberis sempervirens L., Iberis umbellata L. in Peninsular Ranges, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern Outer North Coast Ranges, Central Coast. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

NOCCAEA
[Biennial] perennial herb, generally glaucous; hairs simple or 0. Stem: erect or decumbent. Leaf: basal rosetted, petioled, entire or dentate; cauline sessile, base lobed or clasping. Inflorescence: elongated or not. Flower: sepals erect, lateral pair not sac-like at base; petals white to ± purple, obscurely clawed. Fruit: silicle, flat perpendicular to septum, dehiscent, unsegmented; valves keeled, generally winged; stigma entire. Seed: 2–10[24], in 1 row, wingless.
80 species: temperate, generally northern hemisphere. (D. Nocca, botanist, director of botanical garden, Pavia, Italy 1758–1841) [Koch & Al-Shehbaz 2004 Syst Bot 29:375–384]
Unabridged references: [Holmgren 1971 Mem New York Bot Gard 21:1–106]

N. fendleri (A. Gray) Holub
NATIVE
Plant ± glaucous; caudex branched or simple. Stem: simple or branched distally. Leaf: petiole 0.4–7.3 cm; blade 0.4–3 cm; cauline ovate to ± oblong. Flower: sepals 1.6–5.3 mm; petals spoon-shaped. Fruit: pedicel 2.5–15 mm. Seed: 2–10, 1.1–2.1 mm, ovate.
2n=28. 5 subspecies. [Online Interchange]
Unabridged synonyms: [Thlaspi fendleri A. Gray; Thlaspi montanum L. var. fendleri (A. Gray) P.K. Holmgren]
Unabridged note: North America plants of this sp. were treated by P.K. Holmgren as conspecific with the European Noccaea montana (as Thlaspi montanum L.), but further studies indicate these are not closely related.

Key to Noccaea fendleri

N. fendleri subsp. californica (S. Watson) Al-Shehbaz & M. Koch KNEELAND PRAIRIE PENNYCRESS
NATIVE
Stem: (5)9.5–11.5(20) cm. Leaf: basal 5–7(8) mm wide, spoon-shaped to obovate, base tapered; petiole 0.8–1.8 × > blade; cauline 2–5. Flower: petals 6–8 mm, 1.6–2.5 mm wide, white. Fruit: 7–10.5 mm, 2.7–4 mm wide, wingless; style (1.3)1.5–2(2.4) mm. Seed: 2–6.
Serpentine outcrops; 500–700 m. Outer North Coast Ranges (Kneeland Prairie). [Thlaspi californicum S. Watson] May–Jun [Online Interchange] {CNPS list}
Unabridged synonyms: [Thlaspi alpestre L. var. californicum (S. Watson) Jeps.; Thlaspi glaucum (A. Nelson) A. Nelson subsp. californicum (S. Watson) Munz; Thlaspi montanum L. var. californicum (S. Watson) P.K. Holmgren]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Jul 30 2014
Citation for this treatment: [Author of taxon treatment] 2013. Noccaea, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=89058, accessed on Jul 30 2014

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Bioregions in which Noccaea fendleri subsp. californica occurs Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.