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PLANTAGINACEAE PLANTAIN FAMILY

Margriet Wetherwax, except as noted

Annual to shrub, some aquatic.
Leaf: basal or cauline, alternate or opposite ( whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0.
Inflorescence: raceme, spike, or flowers axillary in 1–few-flowered clusters; flowers few to many, each subtended by 1 bract.
Flower: unisexual or bisexual, radial or bilateral; sepals 4–5, generally fused at base; corolla 4–5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1–2, anthers opening by 2 slits; ovary superior, [1]2–4-chambered, style 1, stigma lobes 0 or 2.
Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
110 genera, ± 2000 species: worldwide, especially temperate. [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2001 Molec Phylogen Evol 16:96–112] Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic CA genera of Scrophulariaceae (except Buddleja, Limosella, Mimulus, Myoporum, Scrophularia, Verbascum). CA Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated. —Scientific Editors: Robert Patterson, Bruce G. Baldwin.

Key to Plantaginaceae

ANTIRRHINUM SNAPDRAGON

Margriet Wetherwax & David M. Thompson

Annual, perennial herb, glabrous to hairy.
Stem: ascending, erect, or vine-like, often clinging by twining pedicels or branchlets.
Leaf: proximal generally opposite, distal alternate, generally reduced distally on stem; veins pinnate.
Inflorescence: cleistogamous or opening; raceme or flower 1 in axils.
Flower: uppermost calyx lobe generally largest; corolla tube of opening flowers truncate or with rounded sac-like extension at base, lower lip base generally swollen, closing throat; stamens 4, generally included, staminode 0; style included, straight or curved, glabrous or glandular-puberulent to near tip, stigma inconspicuous.
Fruit: ovoid to spheric; chambers 2, generally dehiscent by 1–2 pores near tip, lower chamber generally larger, upper occasionally indehiscent.
Seed: many, generally with tubercles or netted ridges, winged or not.
35 species: w North America, w Medit. (Greek: nose-like, from corolla shape) [Oyama & Baum 2004 Amer J Bot 91:918–925; Vargas et al. 2004 Plant Syst Evol 249:151–172] North America taxa more closely related to Mohavea than to Medit taxa; revision needed. Antirrhinum cyathiferum moved to Pseudorontium.
Unabridged references: [Ghebrehiwet et al. 2000 Plant Syst Evol 220:223–239; Thompson 1988 Syst Bot Monogr 22:1–142]

Key to Antirrhinum

A. vexillocalyculatum Kellogg
NATIVE
Annual, branched or not, glabrous to hairy proximal to inflorescence.
Stem: erect but weak, often clinging to other plants or debris.
Leaf: petiole 0–27 mm; blade 1.5–60 mm, elliptic to ovate or oblong, tip obtuse to rounded.
Inflorescence: raceme-like or flowers 1 in axils, glandular-hairy, flowers all opening; pedicels 1–4 mm, proximal-most subtended by twining branchlets, flower branchlets, or both.
Flower: calyx lobes unequal; corolla lavender, veins often vaguely darker, throat uniformly narrowed, curved upward at mouth, lower lip lobes conspicuous, thrust forward, erect or spreading.
Fruit: 4–8 mm; dehiscent by 3 pores at tip, 1 in upper chamber, 2 in lower.
Seed: 0.7–1.3 mm, ovoid, dark brown to black, tubercled, ridged.
n=15. [Antirrhinum vexillo-calyculatum, orth. var.] Subspp. intergrade in NCoR (except Antirrhinum vexillocalyculatum subsp. vexillocalyculatum). [Online Interchange]

A. vexillocalyculatum subsp. vexillocalyculatum
NATIVE

Stem: 7–170 cm, proximal stem glabrous or hairs sparse, <= 3 mm, nonglandular.
Inflorescence: branchlets subtending flowers with 2 leaves at proximal-most node.
Flower: upper calyx lobe 6.8–14 mm, lower 4–8.6 mm; corolla 11–17 mm, darker veins 0.
Gravelly lower slopes of rockslides, disturbed areas, often on serpentine; < 1200 m. s North Coast Ranges, Sacramento Valley (Sutter Buttes), San Francisco Bay Area, n Inner South Coast Ranges. Jun–Aug [Online Interchange]

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Next taxon: Antirrhinum virga

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Citation for the whole project: Jepson Flora Project (eds.) [year] Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html [accessed on month, day, year]
Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on [month, day, year].

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Bioregions in which taxon occursRed area (if present) is the part of the bioregion lying between the upper and lower elevation limits of the taxon;
markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may have georeferencing or identification issues.
map of distribution 1

Chart based on elevation range in Manual and elevations and coordinates of CCH records.
Data provided by the participants of the Consortium of California Herbaria.
Note: About half of the CCH records include both elevation and coordinates.
Map made in collaboration with Scott Loarie. Data provided by the participants of the Consortium of California Herbaria.
View all CCH records

 

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes Manual flowering time.