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BRASSICACEAE (Cruciferae) MUSTARD FAMILY

Ihsan A. Al-Shehbaz, except as noted

Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1–many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
± 330 genera, 3780 species: worldwide, especially temperate. [Al-Shehbaz et al. 2006 Plant Syst Evol 259:89–120] Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central North Coast Ranges, Carrichtera annua (L.) DC. in South Coast, Iberis sempervirens L., Iberis umbellata L. in Peninsular Ranges, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern Outer North Coast Ranges, Central Coast. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella. —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.

Key to Brassicaceae

STREPTANTHUS JEWELFLOWER
Annual to perennial herb, generally ± glaucous; hairs simple or 0. Leaf: basal rosetted or not, petioled, entire or dentate to pinnately lobed or divided; cauline sessile, occasionally petioled, base generally lobed or clasping. Inflorescence: elongated. Flower: radial or bilateral; calyx urn- or occasionally bell-shaped, sepals erect, base ± sac-like, keeled or not; petal blade narrower to wider than proximal 1/2, generally channeled, margins ± crinkled or not; stamens in 3 pairs of unequal length, or 4 long and 2 short, longest filaments fused or free. Fruit: silique, dehiscent, linear, flat parallel to septum, unsegmented; stigma entire or 2-lobed. Seed: 10–120, in 1 row, generally winged.
35 species: southwestern United States, northern Mexico. (Greek: twisted flower, from wavy-margined petals)

Key to Streptanthus

S. glandulosus Hook.
NATIVE
Annual, stiff-hairy basally (glabrous throughout). Stem: (0.8)1.5–9(12) dm, simple to branched throughout. Leaf: basal not rosetted, petioled, early-deciduous, coarsely dentate to ± lobed; mid-cauline sessile, 1–12 cm, lance-linear to oblanceolate, entire to coarsely dentate, leaf base lobed to clasping; distal similar, reduced, generally entire. Inflorescence: open, 1-sided or not; terminal sterile flower cluster 0. Flower: calyx bilateral, sepals (3)5–10(13) mm, lanceolate to broadly ovate, white to yellow, rose, purple, or purple-black, glabrous or bristles sparse; petals 7–17 mm, 1–3 mm wide, ± equal or adaxial pair longer, crinkled; filaments in 3 pairs of unequal length; longest pair fused, 5–13 mm; fertile anthers 1–2.5 mm. Fruit: ascending to spreading or reflexed, 3–11 cm, 1.5–2.5 mm wide, straight or curved, not constricted between seeds; valve midveins distinct, glabrous or sparsely to moderately hairy; stigma ± entire; pedicels ascending to spreading, 0.2–3.2 cm. Seed: 22–70, 1.5–2.1 mm, ovate to oblong; wing continuous, 0.1–0.5 mm wide. 8 subspecies recognized, 7 in California, most local. [Online Interchange]
Unabridged note: Streptanthus glandulosus subsp. josephinensis endemic to southwestern Oregon.

S. glandulosus subsp. sonomensis (Kruckeb.) M.S. Mayer & D.W. Taylor UNCOMMON JEWELFLOWER
NATIVE
Stem: 2.3–7(9) dm, moderately to densely bristly proximally, glabrous distally. Leaf: cauline bristly; distal entire or finely dentate. Inflorescence: 1-sided, axis straight; pedicels 3–10 mm, sparsely hairy or glabrous. Flower: sepals 5–6 mm, cream or pale yellow, base not ± purple or lavender, glabrous or sparsely hairy; petals 10–17 mm, veins not purple; longest filaments 7–10 mm. Fruit: spreading to ± reflexed, arched, glabrous or hairs sparse.
Serpentine outcrops; ± 300 m. Outer North Coast Ranges (c Sonoma Co.). May–Jul [Online Interchange]
Unabridged synonyms: [Streptanthus glandulosus var. sonomensis Kruckeb.]

Previous taxon: Streptanthus glandulosus subsp. secundus
Next taxon: Streptanthus gracilis

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Dec 19 2014
Citation for this treatment: [Author of taxon treatment] 2013. Streptanthus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=88832, accessed on Dec 19 2014

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Geographic subdivisions indicated for the distribution of Streptanthus glandulosus subsp. sonomensis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.