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Key to families | Table of families and genera
Indexes to all accepted names and synonyms:
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Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, ± fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or ± fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1–3, placentas axile or parietal, ovules many, style generally 1, 2–5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
± 90 genera, ± 2500 species: worldwide. [Haberle et al. 2008 J Molec Evol 66:350–361] Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Lammers 2007 World Checklist and Bibliography of Campanulaceae. Royal Botanic Gardens, Kew.]
Key to Campanulaceae
Annual, from taproot. Stem: prostrate, decumbent, or erect; base generally ± brown or ± purple; branches 0 or below middle. Leaf: basal; petiole short or 0. Inflorescence: ± raceme-like; bract 1 per flower, small; pedicel generally thread-like. Flower: inverted or not; sepals linear to triangular; corolla ± radial or 2-lipped, lobes 5; filaments free at base, fused into tube around style distally, appendages attached to a stalk or directly on 2 adjacent filaments, each with 2–12 cells, anthers free, all alike; ovary superior to 1/2 inferior, hemispheric to obconic, nectary glands 3, mounded or donut-like, on free part of ovary, stigma 2-lobed, papillate. Fruit: generally > hypanthium, hemispheric to fusiform, top pointed or rounded, chambers 2; open at top generally by 2 valves (or circumscissile). Seed: elliptic to oblong.Key to Nemacladus
18 species: southwestern United States, northwestern Mexico. (Greek: thread-like branch) [Morin 2008 J Bot Res Inst Texas 2:397–400] In descriptions, "filaments" including both free and fused parts thereof.
Unabridged references: [McVaugh 1942 N Amer Flora 32A:1–134]
Unabridged note: Taxonomic changes from TJM (1993) based on ITS, atpB, morphology. Parishella californica nested within Nemacladus, in which it is here treated.
Erect, 5–25 cm, branched from base. Stem: stiffly ascending. Leaf: 3–16 mm, oblanceolate to elliptic, toothed or pinnately lobed, hairy, tapered to petiole. Inflorescence: axis ± straight; bracts 1–6 mm, spreading, lanceolate to ovate; pedicels 6–16 mm, 0.1 mm diam, stiffly ascending, straight to tip. Flower: not inverted; hypanthium ± 1 mm; sepals 0.8–2.3 mm, linear-elliptic to ± deltate, erect; corolla 2-lipped, divided ± to base, white, lobes oblanceolate, 3 adaxial erect, 1.3–2.5 mm, maroon at tips, ciliate, 2 abaxial 1.3–2.5 mm; filaments 1–2 mm, declined, tip ± curved, glabrous, appendages stout stalks, cells blunt, anthers 0.2–0.4 mm; ovaries 1/3–1/2-inferior. Fruit: ± 2–4 mm, base rounded, tip acute. Seed: ± 0.5 mm, cylindric, with impressed, vertical lines crossed by fine transverse lines.
Dry slopes, sandy soils, washes; < 2400 m. Inner South Coast Ranges, San Gabriel Mountains, n San Bernardino Mountains, East of Sierra Nevada, Desert; to Utah, New Mexico, Baja California. [Nemacladus glanduliferus Jeps. var. orientalis McVaugh] Mar–May [Online Interchange]
Previous taxon: Nemacladus montanus
Next taxon: Nemacladus pinnatifidus
Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Jul 27 2015
Citation for this treatment: [Author of taxon treatment] 2013. Nemacladus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=88823, accessed on Jul 27 2015
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© 2010 James Morefield
|Geographic subdivisions indicated for the distribution of Nemacladus orientalis|| Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
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