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RHAMNACEAE BUCKTHORN FAMILY

John O. Sawyer, Jr., except as noted

[Perennial herb] shrub, tree, generally erect, often thorny. Leaf: simple, generally alternate, often clustered on short-shoots; stipules generally present, occasionally modified into spines; generally petioled; blade pinnate-veined or 1–5-ribbed from base. Inflorescence: cyme, panicle, umbel, or flowers 1 or clustered in axils or on short-shoots. Flower: generally bisexual, radial; hypanthium subtending, surrounding, or partly fused to ovary; sepals 4–5; petals 0, 4–5, generally clawed; stamens 0, 4–5, alternate sepals, attached to hypanthium top, each generally fitting into a petal concavity; disk (0 or) between stamens, ovary, thin to fleshy, entire or lobed, free from ovary, adherent or fused to hypanthium; ovary superior or ± inferior, chambers [1]2–4, 1–2-ovuled, style 1, stigma entire or 2–3-lobed. Fruit: capsule, drupe.
50–52 genera, 950 species: especially tropics, subtropics some cultivated (Ceanothus; Frangula; Rhamnus; Ziziphus). [Richardson et al. 2000 Amer J Bot 87:1309–1324] —Scientific Editors: Steve Boyd, Thomas J. Rosatti.
Unabridged references: [Brizicky 1965 J Arnold Arbor 45:439–463; Richardson et al. 2000 Kew Bull 55:311–341]

Key to Rhamnaceae

CONDALIA
Shrub. Stem: branches alternate, rigid; twigs thorn-tipped. Leaf: clustered on short-shoots, deciduous; stipules deciduous; petioles ± 0; blade generally obovate, 1-ribbed from base, entire. Inflorescence: flowers 1 or in clusters on short-shoots. Flower: hypanthium hemispheric, 1–1.5 mm wide; sepals 5, deciduous; petals 0 [5]; stamens 5; disk 0 or thin in early flower; ovary spheric, strong-narrowed at base, chambers 2, each 1-ovuled, stigma entire. Fruit: drupe, stone 1.
18 species: arid America. (A. Condal, Spanish physician, 1745?–1804) [Christie et al. 2006 Canotia 2:23–46]
Unabridged references: [Johnston 1962 Brittonia 14:332–368]

Previous taxon: Colubrina californica
Next taxon: Condalia globosa var. pubescens

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 21 2014
Citation for this treatment: [Author of taxon treatment] 2013. Condalia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=8836, accessed on Aug 21 2014

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