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RHAMNACEAE BUCKTHORN FAMILY

John O. Sawyer, Jr., except as noted

[Perennial herb] shrub, tree, generally erect, often thorny. Leaf: simple, generally alternate, often clustered on short-shoots; stipules generally present, occasionally modified into spines; generally petioled; blade pinnate-veined or 1–5-ribbed from base. Inflorescence: cyme, panicle, umbel, or flowers 1 or clustered in axils or on short-shoots. Flower: generally bisexual, radial; hypanthium subtending, surrounding, or partly fused to ovary; sepals 4–5; petals 0, 4–5, generally clawed; stamens 0, 4–5, alternate sepals, attached to hypanthium top, each generally fitting into a petal concavity; disk (0 or) between stamens, ovary, thin to fleshy, entire or lobed, free from ovary, adherent or fused to hypanthium; ovary superior or ± inferior, chambers [1]2–4, 1–2-ovuled, style 1, stigma entire or 2–3-lobed. Fruit: capsule, drupe.
50–52 genera, 950 species: especially tropics, subtropics some cultivated (Ceanothus; Frangula; Rhamnus; Ziziphus). [Richardson et al. 2000 Amer J Bot 87:1309–1324] —Scientific Editors: Steve Boyd, Thomas J. Rosatti.
Unabridged references: [Brizicky 1965 J Arnold Arbor 45:439–463; Richardson et al. 2000 Kew Bull 55:311–341]

Key to Rhamnaceae

ADOLPHIA
Shrub. Stem: branches dense, opposite, rigid, green; twigs spreading, jointed at base, thorn-tipped, glabrous to puberulent. Leaf: opposite, clustered in axils or not, early-deciduous; stipules ± triangular, black, shiny; blade entire, 1–3-ribbed from base. Inflorescence: small axillary clusters; pedicel generally white to green-white. Flower: hypanthium bowl-shaped, generally white to green-white; sepals 5, triangular, persistent; petals 5, spoon-shaped, white; stamens 5; disk fused to hypanthium, 5-angled, fleshy, generally white to green-white; ovary superior, chambers 3, 1-ovuled, stigma 3-lobed. Fruit: capsule, spheric, explosively dehiscent.
2 species: California, Texas, Mexico. (Adolphe T. Brongniart, student of Rhamnaceae, 1801–1876)

Previous taxon: Rhamnaceae
Next taxon: Adolphia californica

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Dec 22 2014
Citation for this treatment: [Author of taxon treatment] 2013. Adolphia, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=8830, accessed on Dec 22 2014

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