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NYCTAGINACEAE FOUR O'CLOCK FAMILY

Andy Murdock, except as noted

Annual, perennial herb, subshrub, [shrub, tree], glabrous or hairy. Stem: often forked. Leaf: opposite, sessile or petioled, pairs generally unequal; blade generally entire. Inflorescence: generally forked; spike, head-like cluster, or umbel, flowers rarely 1, bracts forming a calyx-like involucre or not. Flower: bisexual, generally ± radial (bilateral), sometimes cleistogamous in some genera; perianth of 1 whorl, generally petal-like, bell- to trumpet-shaped, base hardened, tightly surrounding ovary in fruit, lobes 3–5, generally notched to ± lobed; stamens 1–many; ovary superior (appearing inferior due to hardened perianth base), style 1. Fruit: achene in hardened perianth base; round to ± flat; smooth, angled, ribbed, or winged; glabrous, hairy, or glandular.
30 genera, 350 species: warm regions, especially America; some ornamental (Bougainvillea; Mirabilis, four o'clock). [Spellenberg 2003 FNANM 4:14–17] —Scientific Editor: Thomas J. Rosatti.

Key to Nyctaginaceae

MIRABILIS FOUR O'CLOCK
Perennial herb, subshrub. Stem: repeatedly forked, decumbent to erect. Leaf: generally petioled. Inflorescence: branches ending in umbel-like cluster or solitary flowers; bracts 5(9), ± fused (or not) into calyx-like, bell- to saucer-shaped involucre; flowers in 1 involucre 1–16, generally not blooming together; flowers cleistogamous or not. Flower: radial or ± bilateral; perianth funnel- to bell-shaped, lobes 5; stamens 3–5, generally exserted; stigma ± spheric, generally exserted. Fruit: ± round to club-shaped; ribs or angles 0, 5, 10; wings 0.
± 60 species: America, Himalayas. (Latin: wonderful) [Spellenberg 2003 FNANM 4:40–57] Flowers open in evening, close in morning; species intergrade, taxonomy unsettled.

Key to Mirabilis

Previous taxon: Boerhavia wrightii
Next taxon: Mirabilis albida

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Oct 20 2014
Citation for this treatment: [Author of taxon treatment] 2013. Mirabilis, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=8789, accessed on Oct 20 2014

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