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NYCTAGINACEAE FOUR O'CLOCK FAMILY

Andy Murdock, except as noted

Annual, perennial herb, subshrub, [shrub, tree], glabrous or hairy. Stem: often forked. Leaf: opposite, sessile or petioled, pairs generally unequal; blade generally entire. Inflorescence: generally forked; spike, head-like cluster, or umbel, flowers rarely 1, bracts forming a calyx-like involucre or not. Flower: bisexual, generally ± radial (bilateral), sometimes cleistogamous in some genera; perianth of 1 whorl, generally petal-like, bell- to trumpet-shaped, base hardened, tightly surrounding ovary in fruit, lobes 3–5, generally notched to ± lobed; stamens 1–many; ovary superior (appearing inferior due to hardened perianth base), style 1. Fruit: achene in hardened perianth base; round to ± flat; smooth, angled, ribbed, or winged; glabrous, hairy, or glandular.
30 genera, 350 species: warm regions, especially America; some ornamental (Bougainvillea; Mirabilis, four o'clock). [Spellenberg 2003 FNANM 4:14–17] —Scientific Editor: Thomas J. Rosatti.

Key to Nyctaginaceae

ACLEISANTHES TRUMPETS
Perennial herb from thick taproot, minutely hairy. Stem: prostrate to ascending. Leaf: petioled; blade < 8 cm, ± firm to fleshy, paired leaves ± unequal. Inflorescence: bracts 1–3, free, not forming involucre; flower generally 1, or paired. Flower: nocturnal or cleistogamous; perianth trumpet-shaped, white, tube slender; stamens 5, ± exserted; stigma ± spheric, ± exceeding anthers. Fruit: oblong, glabrous; wings translucent or 0.
17 species: especially southwestern United States, northeastern Mexico. (Greek: without closure, from absence of involucre) [Poole 2003 FNANM 4:33–39] Pollinated by hawkmoths. Selinocarpus moved to Acleisanthes.

Key to Acleisanthes

Previous taxon: Abronia villosa var. villosa
Next taxon: Acleisanthes longiflora

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Aug 30 2014
Citation for this treatment: [Author of taxon treatment] 2013. Acleisanthes, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=8781, accessed on Aug 30 2014

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