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ASTERACEAE (Compositae) SUNFLOWER FAMILY

David J. Keil, except as noted

Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1–many in generally ± cyme-like cluster; each head generally with ± calyx-like involucre of 1–many series of phyllaries (involucral bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1–many per head. Flower: bisexual, unisexual, or sterile, ± small, of several types (see below); calyx 0 or modified into ± persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3–5; stamens 4–5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally ± 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing ± brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
± 1500 genera, 23000 species: worldwide, many habitats. Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Eucephalus, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Bahia; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Taxa of Arida in TJM2 treated here in Leucosyris. —Scientific Editors: David J. Keil, Bruce G. Baldwin.
Unabridged note: Largest family of vascular plants in California and of eudicots globally.

Key to Asteraceae

VOLUTARIA
Annual. Stem: erect, openly branched. Leaf: basal and proximal cauline winged-petioled, mid and distal cauline sessile; entire to dentate or pinnately divided, long-soft hairy, minutely glandular. Inflorescence: heads radiant, long-peduncled, 1 or in few-headed cyme-like cluster; involucre ovoid, 10–15 mm diam, phyllaries many, graduated in several series, appressed, ovate to lanceolate, entire, acute, tipped by ascending to reflexed, flattened spine; receptacle flat, epaleate, bristly. Flower: corolla pink to purple [blue, yellow, or orange]; peripheral flowers sterile, corolla enlarged, spreading, lobes (5–6), linear; inner flowers bisexual, tube slender, throat narrowly cylindric, lobes linear-oblong; anther bases tailed, tips lanceolate; style tip with ± hairy node and short, linear terminal segment. Fruit: ± barrel-shaped, weakly compressed, ribbed, tip with prominent collar, faces pitted, attachment scar lateral within cavity with hard rim; pappus persistent, of several series of narrow scales, white to tan.
16 species: Mediterranean, western Asia, Atlantic islands, northeastern Africa. (Latin: having twist, for spirally coiled corolla lobes of original sp.) [Keil 2006 FNANM 19:174–175] Volutaria canariensis Wagenitz (disciform heads, white flowers) established as weed in southwestern Sonoran Desert.
Unabridged synonyms: [Cyanopsis]

V. muricata (L.) Maire MOROCCO KNAPWEED
NATURALIZED
Stem: <= 50 cm. Leaf: proximal oblanceolate, ± dentate to deeply lobed, distal smaller, linear, ± entire or lobed, long-shaggy-hairy (hairs jointed), minutely resin-gland-dotted. Inflorescence: head 3–5 cm diam; peduncle 5–15 cm, distally leafless, finely cobwebby-tomentose, especially near tips; involucre 15–18 mm, phyllaries in 5–7 series, minutely spiny-serrate, finely soft hairy, outer and middle ovate, dark-margined, with spine tip 3–5 mm, inner lanceolate, with flattened, ± spineless tips. Flower: sterile flower corolla lobes 10–15 mm, spreading; bisexual flower corolla tube 5–6 mm, throat 2–4 mm, lobes 5–6 mm. Fruit: pale gray-brown; pappus scales 1–2.5 mm, irregularly toothed, outer < inner.
2n=24. Disturbed sites; < 100 m. South Coast; southwestern Europe, northwestern Africa. [Centaurea muricata L.; Cyanopsis muricata (L.) Dostál] Probable escape from cultivation; becoming problem weed in southern South Coast. Mar–May [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Sep 2 2014
Citation for this treatment: [Author of taxon treatment] 2013. Volutaria, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=82516, accessed on Sep 2 2014

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Geographic subdivisions indicated for the distribution of Volutaria muricata Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.