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Daniel Potter & Barbara Ertter, family description, key to genera; treatment of genera by Daniel Potter, except as noted

Annual to tree, glandular or not. Leaf: simple to palmately or pinnately compound, generally alternate; stipules free to fused (0), persistent to deciduous. Inflorescence: cyme, raceme, panicle, cluster, or flowers 1; bractlets on pedicel ("pedicel bractlets") generally 0–3(many), subtended by bract or generally not. Flower: generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, subtending bractlets ("hypanthium bractlets") 0–5, alternate sepals; sepals generally 5; petals generally 5, free; stamens (0,1)5–many, anther pollen sacs generally 2; pistils (0)1–many, simple or compound, ovary superior to inferior, styles 1–5. Fruit: 1–many per flower, achene (fleshy-coated or not), follicle, drupe, or pome with generally papery core, occasionally drupe-like with 1–5 stones. Seed: generally 1–5 (per fruit, not per flower).
110 genera, ± 3000 species: worldwide, especially temperate; many cultivated for ornamental, fruit, especially Cotoneaster, Fragaria, Malus, Prunus, Pyracantha, Rosa, Rubus. [Potter et al. 2007 Plant Syst Evol 266:5–43] Number of teeth is per leaf or leaflet, not per side of leaf or leaflet, except in Drymocallis. —Scientific Editors: Daniel Potter, Thomas J. Rosatti.
Unabridged references: [Robertson 1974 J Arnold Arbor 55:303–332, 344–401, 611–662]

Key to Rosaceae


Barbara Ertter

Perennial herb, hairs short, simple, nonglandular and/or long, cross-walled, glandular. Stem: ± erect, from ± branched caudex or rhizomes. Leaf: basal and cauline, alternate, odd-1-pinnately compound; leaflets toothed, terminal generally >= lateral. Inflorescence: cyme, ± open; pedicels straight, bractlets 0. Flower: hypanthium ± shallow, bractlets 5; sepals ± triangular; petals < to > sepals, white to yellow; stamens generally 20–25, pollen sac 1, horseshoe-shaped; pistils many, styles fusiform, attached below fruit middle. Fruit: achene, glabrous.
30 species: northern temperate. (Greek: wood beauty) [Ertter 2007 J Bot Res Inst Texas 1:31–46] Recognition based on morphological, molecular evidence. Drymocallis ashlandica (Green) Rydb. (inflorescence narrow, petioles glandular, petals yellow) in southwestern Oregon, possibly northwestern California.
Unabridged note: Convergence of morphological, molecular evidence mandates recognition of Drymocallis; infrageneric taxonomy provisional. Key characters include relative proportions of 2 hair types: short simple nonglandular hairs and longer cross-walled glandular hairs.

Key to Drymocallis

D. glandulosa (Lindl.) Rydb.
Tufted. Stem: glandular hairs abundant at base. Leaf: sheathing base glabrous to glandular. Inflorescence: pedicel glandular hairs generally abundant, short nonglandular hairs generally sparse. Flower: opening widely; hypanthium bractlets ± elliptic; petals spreading; styles generally ± 1 mm. Fruit: 1–1.5 mm, ± red to brown. [Potentilla glandulosa Lindl.] [Online Interchange]

D. glandulosa var. reflexa (Greene) Ertter
Stem: generally 15–80 cm. Leaf: basal generally 6–15 cm, lateral leaflet pairs generally 3, terminal leaflet generally 15–40 mm, obovate, teeth ± double, ± 6–16 per side. Inflorescence: not leafy, spreading, branch angle generally 25–55°; pedicels generally 2–10 mm, lowermost to 20 mm. Flower: hypanthium bractlets 1.5–4 mm, 0.5–1 mm wide; sepals ± reflexed, generally 4–6 mm, ± obtuse; petals generally 3–4 mm, 2–2.5 mm wide, narrow-obovate, yellow.
Moist or ± shaded places; 450–2600 m. Klamath Ranges, High North Coast Ranges, High Cascade Range, High Sierra Nevada, San Gabriel Mountains, San Bernardino Mountains, Peninsular Ranges; southern Oregon, western Nevada, northern Baja California. [Potentilla glandulosa subsp. reflexa (Greene) D.D. Keck] May–Aug [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Mar 30 2015
Citation for this treatment: [Author of taxon treatment] 2013. Drymocallis, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Mar 30 2015

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Geographic subdivisions indicated for the distribution of Drymocallis glandulosa var. reflexa Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.