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Daniel Potter & Barbara Ertter, family description, key to genera; treatment of genera by Daniel Potter, except as noted

Annual to tree, glandular or not. Leaf: simple to palmately or pinnately compound, generally alternate; stipules free to fused (0), persistent to deciduous. Inflorescence: cyme, raceme, panicle, cluster, or flowers 1; bractlets on pedicel ("pedicel bractlets") generally 0–3(many), subtended by bract or generally not. Flower: generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, subtending bractlets ("hypanthium bractlets") 0–5, alternate sepals; sepals generally 5; petals generally 5, free; stamens (0,1)5–many, anther pollen sacs generally 2; pistils (0)1–many, simple or compound, ovary superior to inferior, styles 1–5. Fruit: 1–many per flower, achene (fleshy-coated or not), follicle, drupe, or pome with generally papery core, occasionally drupe-like with 1–5 stones. Seed: generally 1–5 (per fruit, not per flower).
110 genera, ± 3000 species: worldwide, especially temperate; many cultivated for ornamental, fruit, especially Cotoneaster, Fragaria, Malus, Prunus, Pyracantha, Rosa, Rubus. [Potter et al. 2007 Plant Syst Evol 266:5–43] Number of teeth is per leaf or leaflet, not per side of leaf or leaflet, except in Drymocallis. —Scientific Editors: Daniel Potter, Thomas J. Rosatti.
Unabridged references: [Robertson 1974 J Arnold Arbor 55:303–332, 344–401, 611–662]

Key to Rosaceae


Barbara Ertter

Perennial herb, hairs short, simple, nonglandular and/or long, cross-walled, glandular. Stem: ± erect, from ± branched caudex or rhizomes. Leaf: basal and cauline, alternate, odd-1-pinnately compound; leaflets toothed, terminal generally >= lateral. Inflorescence: cyme, ± open; pedicels straight, bractlets 0. Flower: hypanthium ± shallow, bractlets 5; sepals ± triangular; petals < to > sepals, white to yellow; stamens generally 20–25, pollen sac 1, horseshoe-shaped; pistils many, styles fusiform, attached below fruit middle. Fruit: achene, glabrous.
30 species: northern temperate. (Greek: wood beauty) [Ertter 2007 J Bot Res Inst Texas 1:31–46] Recognition based on morphological, molecular evidence. Drymocallis ashlandica (Green) Rydb. (inflorescence narrow, petioles glandular, petals yellow) in southwestern Oregon, possibly northwestern California.
Unabridged note: Convergence of morphological, molecular evidence mandates recognition of Drymocallis; infrageneric taxonomy provisional. Key characters include relative proportions of 2 hair types: short simple nonglandular hairs and longer cross-walled glandular hairs.

Key to Drymocallis

D. pseudorupestris (Rydb.) Rydb.
Tufted to matted. Stem: generally 5–25 cm, glandular hairs abundant at base. Leaf: basal generally 6–9 cm, sheathing base generally appressed-hairy, terminal leaflet generally 5–20 mm, widely obovate to fan-shaped, ± rounded, teeth generally ± single, 4–10 per side. Inflorescence: not leafy, spreading, branch angle generally 20–40°; pedicels generally 3–10 mm, lowermost to 20 mm, glandular hairs generally abundant, short nonglandular hairs 0–many. Flower: opening widely; hypanthium bractlets 2–5 mm, 1–2 mm wide, lance-linear to elliptic-ovate; sepals generally 4–6 mm, acute to obtuse; petals spreading, generally 4–8 mm, > sepals, ± obovate, cream to pale yellow; styles 1–1.5 mm. Fruit: ± 1 mm, light brown. [Potentilla glandulosa subsp. pseudorupestris (Rydb.) D.D. Keck] [Online Interchange]

D. pseudorupestris var. saxicola Ertter CLIFF WOODBEAUTY
Stem: short nonglandular hairs generally present. Leaf: leaflets generally 3 per side. Flower: calyx with nonglandular hairs < 1 mm or 0; styles generally golden-brown.
Rocky areas; 2300–3500 m. Northwestern California, High Cascade Range, High Sierra Nevada, Warner Mountains, White and Inyo Mountains; to British Columbia, Montana. Jul–Sep [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 28 2015
Citation for this treatment: [Author of taxon treatment] 2013. Drymocallis, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 28 2015

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Geographic subdivisions indicated for the distribution of Drymocallis pseudorupestris var. saxicola Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.