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Daniel Potter & Barbara Ertter, family description, key to genera; treatment of genera by Daniel Potter, except as noted

Annual to tree, glandular or not. Leaf: simple to palmately or pinnately compound, generally alternate; stipules free to fused (0), persistent to deciduous. Inflorescence: cyme, raceme, panicle, cluster, or flowers 1; bractlets on pedicel ("pedicel bractlets") generally 0–3(many), subtended by bract or generally not. Flower: generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, subtending bractlets ("hypanthium bractlets") 0–5, alternate sepals; sepals generally 5; petals generally 5, free; stamens (0,1)5–many, anther pollen sacs generally 2; pistils (0)1–many, simple or compound, ovary superior to inferior, styles 1–5. Fruit: 1–many per flower, achene (fleshy-coated or not), follicle, drupe, or pome with generally papery core, occasionally drupe-like with 1–5 stones. Seed: generally 1–5 (per fruit, not per flower).
110 genera, ± 3000 species: worldwide, especially temperate; many cultivated for ornamental, fruit, especially Cotoneaster, Fragaria, Malus, Prunus, Pyracantha, Rosa, Rubus. [Potter et al. 2007 Plant Syst Evol 266:5–43] Number of teeth is per leaf or leaflet, not per side of leaf or leaflet, except in Drymocallis. —Scientific Editors: Daniel Potter, Thomas J. Rosatti.
Unabridged references: [Robertson 1974 J Arnold Arbor 55:303–332, 344–401, 611–662]

Key to Rosaceae


Barbara Ertter

Perennial herb, tufted, ± glandular; odor ± resinous. Stem: ascending to erect, 15–50(90) cm. Leaf: generally basal, odd-pinnately compound; leaflets 15–35 per side, ± overlapped, toothed to palmately divided, segments ± oblanceolate. Inflorescence: ± cyme; pedicels straight, bractlets 0. Flower: hypanthium cup-like, flat-bottomed, bractlets 5; sepals often reflexed; petals white or ± pink, midvein often ± red; stamens 20, filaments ± flat, often an erect tube; pistils many, ovaries superior, style attached below fruit tip, ± rough, thick. Fruit: achene, ± 1.5 mm.
2 species: California. (Latin: small Horkelia) Flower ± like Horkelia.

Key to Horkeliella

H. congdonis (Rydb.) Rydb.
Plant green, generally glandular-sticky. Leaf: basal generally 8–25 cm, leaflets 25–35 per side, generally 2–6 mm; cauline 2–7. Inflorescence: ± dense; pedicels generally ± 2 mm. Flower: ± 15 mm wide; sepals 4–8 mm; petals 3–6 mm, oblong-oblanceolate; anthers ± 1 mm, filaments 1.5–3 mm; style 2–5 mm.
Meadows in sagebrush flats; 1500–3100 m. c High Sierra Nevada (e slope), w East of Sierra Nevada, Desert Mountains (Coso Range). [Ivesia purpurascens (S. Watson) D.D. Keck subsp. congdonis (Rydb.) D.D. Keck] Jul–Aug [Online Interchange]

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 28 2015
Citation for this treatment: [Author of taxon treatment] 2013. Horkeliella, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 28 2015

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Geographic subdivisions indicated for the distribution of Horkeliella congdonis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.